2011 in paleoichthyology

Fishes

Newly named jawless vertebrates

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Althaspis tarloi[1]	sp.	Valid	Voichyshyn	Early Devonian	Dniester Formation	Ukraine	A heterostracan
Djurinaspis secunda[1]	sp.	Valid	Voichyshyn	Early Devonian	Tyver Formation	Ukraine	A heterostracan
Lechriaspis[2]	Gen. et sp.	Valid	Elliott Petriello	Early Devonian	Water Canyon Formation	USA	A heterostracan
Palanasaspis[1]	Gen. et sp.	Valid	Voichyshyn	Early Devonian	Dniester Formation	Ukraine	A heterostracan
Podolaspis danieli[1]	sp.	Valid	Voichyshyn	Early Devonian	Dniester Formation	Ukraine	A heterostracan
Poraspis thomasi[2]	Species	Valid	Elliott Petriello	Early Devonian	Lost Burro Formation	USA	A heterostracan
Semipodolaspis[1]	Gen. et sp.	Valid	Voichyshyn	Early Devonian	Tyver Formation	Ukraine	A heterostracan
Shuyu[3]	Gen. et comb.	Valid	Gai Donoghue et al.[CAL 1]	Late Telychian/Early Wenlock	Maoshan Formation	China	The basalmost known eugaleaspidiform galeaspid.
Zenaspis kasymyri[1]	sp.	Valid	Voichyshyn	Early Devonian	Dniester Formation	Ukraine	A osteostracan

Newly named acanthodian

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Nerepisacanthus[4]	Gen. et sp.	Valid	Burrow	?Telychian/Ludlow	Cunningham Creek Formation	Canada	An acritolepid ischnacanthiform.

Newly named placoderms

Name	Status	Authors	Age	Unit	Location	Notes	Images
Arabosteus[5]	Valid	Olive et al.	Early Devonian	Jauf Formation	Saudi Arabia	A member of Acanthothoraci. The type species is Arabosteus variabilis.
Draconichthys elegans[6]	Valid	Rücklin	Frasnian	Anti-Atlas Mountains	Morocco	A piscivorous selenosteid arthrodire

Newly named cartilaginous fishes

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Cederstroemia ziaensis[7]	Species	Valid	Bourdon et al.	Late Cretaceous (Santonian)	Point Lookout Sandstone	United States	A carpet shark, a species of Cederstroemia.
Dwardius sudindicus[8]	Species	Valid	Underwood Goswami et al.[CAL 2]	Early Cenomanian	Karai Formation	India	An otodontid.
Eorhinoptera[9]	Gen. et sp. nov	Valid	Case, Cook & Wilson	Early Eocene (Ypresian)	Fishburne Formation	United States	A stingray related to the genus Rhinoptera. The type species is Eorhinoptera grabdai.
Ertychius[10]	Gen. et sp. nov	Valid	Ginter, Hairapetian & Grigoryan	Devonian (early Famennian)		Armenia	A ctenacanthiform. The type species is Ertychius intermedius.
Fayolia sharovi[11]	Species	Valid	Fischer Voigt et al.[CAL 3]	Middle/Late Triassic	Madygen Formation	Kyrgyzstan	Egg capsules of a probable xenacanthid.
Gladioserratus[8]	Gen. et comb. et sp. nov	Valid	Underwood Goswami et al.	Cretaceous (Hauterivian) to Paleocene (Danian) or possibly even early Eocene[12]		Australia  Denmark[12]  France  India  United Kingdom	A cow shark, a new genus for "Notidanus" aptiensis (Pictet, 1865). Genus also contains a new species Gladioserratus magnus; it might also contain Eocene species "Notorynchus" serratissimus Agassiz (1843).[12]
Karksiodus[13]	Gen. et sp. nov	Valid	Ivanov, Märss & Kleesment	Devonian (Givetian)		Estonia	A member of Elasmobranchii of uncertain phylogenetic placement. The type species is Karksiodus mirus.
Kawichthys[14]	Gen. et sp.	Valid	Pradel Tafforeau et al.[CAL 4]	Late Virgilian	Douglas Group	USA	A symmoriiform.
Lissodus lusavorichi[10]	Sp. nov	Valid	Ginter, Hairapetian & Grigoryan	Devonian (late Famennian)		Armenia  Iran	A lonchidiid hybodontiform, a species of Lissodus.
Lonchidion ferganensis[11]	Species	Valid	Fischer Voigt et al.	Middle/Late Triassic	Madygen Formation	Kyrgyzstan	A hybodontid shark.
Moerigaleus[15]	Gen. et sp. nov	Valid	Underwood & Ward	Late Eocene		Egypt	A hemigaleid. The type species is Moerigaleus vitreodon.
Palaeoxyris alterna[11]	Species	Valid	Fischer Voigt et al.	Middle/Late Triassic	Madygen Formation	Kyrgyzstan	A hybodontid shark.
Ptychodus atcoensis[16]	Species	Valid	Hamm Cicimurri	Late Cretaceous (early Coniacian)	Atco Formation	United States	A species of Ptychodus.
Pueblocarcharias[7]	Gen. et sp. nov	Valid	Bourdon et al.	Late Cretaceous (Santonian)	Point Lookout Sandstone	United States	A sand shark. The type species is Pueblocarcharias kawaikensis.
Reginaselache[17]	Gen. et sp.	Valid	Turner Burrow	Middle Viséan	Ducabrook Formation	Australia	A xenacanthiform.
Scapanorhynchus puercoensis[7]	Sp. nov	Valid	Bourdon et al.	Late Cretaceous (Santonian)	Point Lookout Sandstone	United States	A mitsukurinid, a species of Scapanorhynchus.
Scindocorax[7]	Gen. et sp. nov	Valid	Bourdon et al.	Late Cretaceous (Santonian)	Point Lookout Sandstone	United States	An anacoracid. The type species is Scindocorax novimexicanus.
Tassiliodus[18]	Gen. et sp.	Valid	Derycke Goujet	Emsian, Devonian		Algeria	An euselachian.
Texatrygon stouti[7]	Sp. nov	Valid	Bourdon et al.	Late Cretaceous (Santonian)	Point Lookout Sandstone	United States	A relative of Ptychotrygon and sclerorhynchids, a species of Texatrygon.

Newly named bony fishes

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Apogon dominans[19]	Sp. nov	Valid	Brzobohatý Nolf	Eocene (Bartonian)		Spain	A cardinalfish, a species of Apogon.
"genus aff. Apogon" viaboadai[19]	Sp. nov	Valid	Brzobohatý Nolf	Eocene (Bartonian)		Spain	A cardinalfish related to members of the genus Apogon.
Apricenaichthys[20]	Gen. et sp. nov	Valid	Taverne	Late Cretaceous (Santonian)		Italy	A beardfish. The type species is Apricenaichthys italicus.
Araripichthys weberi[21]	Species	Valid	Alvarado-Ortega Brito	Albian	Tlayúa Formation	Mexico	An elopocephalan, a species of Araripichthys.
Arius subtilis[22]	Sp. nov	Valid	Schwarzhans Bratishko	Paleocene (Selandian)		Ukraine	An ariid catfish, a species of Arius.
Astroscopus countermani[23]	Species	Valid	Carnevale, Godfrey & Pietsch	Miocene (Tortonian)	St. Marys Formation	United States	A stargazer, a species of Astroscopus.
Aulorhamphus chiarasorbiniae[24]	Sp. nov	Valid	Tyler & Bannikov	Eocene	Monte Bolca locality	Italy	An aulorhamphid gasterosteiform, a species of Aulorhamphus.
Beishanichthys[25]	Gen. et sp.	Valid	Xu Gao	Early Triassic		China	A scanilepiform.
Belonostomus lamarquensis[26]	Sp. nov	Valid	Bogan Taverne Agnolin	Late Cretaceous (Campanian-Maastrichtian)	Allen Formation	Argentina	An aspidorhynchiform.
Bourbonnella jocelynae[27]	Species	Valid	Mickle	Carboniferous	Manning Canyon Shale Formation	USA	An aeduellid.
"genus Bythitidarum" rozenbergi[22]	Sp. nov	Valid	Schwarzhans Bratishko	Paleocene (Selandian)		Ukraine	A brotula of uncertain generic assignment.
Caucasichthys[28]	Gen. et sp.	Valid	Bannikov Carnevale Parin	Bartonian		Russia	A perciform.
Centroberyx anguinicauda[22]	Sp. nov	Valid	Schwarzhans Bratishko	Paleocene (Selandian)		Ukraine	A member of Berycidae, a species of Centroberyx.
Chaetodipterus steurbauti[19]	Sp. nov	Valid	Brzobohatý Nolf	Eocene (Bartonian)		Spain	A spadefish, a species of Chaetodipterus.
Chlorophthalmus udovichenkoi[22]	Sp. nov	Valid	Schwarzhans Bratishko	Paleocene (Selandian)		Ukraine	A greeneye, a species of Chlorophthalmus.
Dapedium stollorum[29]	Species	Valid	Thies Hauff	Early Toarcian		Germany	A species of Dapedium.
"genus Epigonidarum" tyassminensis[22]	Sp. nov	Valid	Schwarzhans Bratishko	Paleocene (Selandian)		Ukraine	A deepwater cardinalfish of uncertain generic assignment.
Fierasferoides bucculentus[22]	Sp. nov	Valid	Schwarzhans Bratishko	Paleocene (Selandian)		Ukraine	A pearlfish, a species of Fierasferoides.
Gadophycis serratus[22]	Sp. nov	Valid	Schwarzhans Bratishko	Paleocene (Selandian)		Ukraine	A cusk-eel, a species of Gadophycis.
Guntherichthys[27]	Gen. et sp.	Valid	Mickle	Carboniferous	Manning Canyon Shale Formation	USA	A ’palaeoniscoid’.
Gwawinapterus	Gen. et sp.	Valid	Arbour Currie	Campanian	Northumberland Formation	Canada	Initially thought to be an istiodactylid pterosaur,[30] but subsequently reinterpreted as an indeterminate saurodontid fish.[31]
"genus Haemulidarum" makarenkoi[22]	Sp. nov	Valid	Schwarzhans Bratishko	Paleocene (Selandian)		Ukraine	A grunt of uncertain generic assignment.
Heteroconger astroblematicus[22]	Sp. nov	Valid	Schwarzhans Bratishko	Paleocene (Selandian)		Ukraine	A member of Congridae, a species of Heteroconger.
"genus Holocentridarum" ryabchuni[22]	Sp. nov	Valid	Schwarzhans Bratishko	Paleocene (Selandian)		Ukraine	A member of Holocentridae of uncertain generic assignment.
Laccognathus embryi[32]	Species	Valid	Downs Daeschler et al.[CAL 5]	Frasnian	Fram Formation	Canada	A porolepiform sarcopterygian.
Lecceclupea[33]	Gen. et sp. nov	Valid	Taverne	Late Cretaceous (Campanian-Maastrichtian boundary)		Italy	A member of Clupeidae belonging to the subfamily Pellonulinae and the tribe Ehiravini. The type species is Lecceclupea ehiravaensis.
"genus Leiognathidarum" tashlikensis[22]	Sp. nov	Valid	Schwarzhans Bratishko	Paleocene (Selandian)		Ukraine	A ponyfish of uncertain generic assignment.
Lepidotes pankowskii[34]	Species	Valid	Forey López-Arbarello MacLeod	Cenomanian	Kem Kem Beds	Morocco	A species of Lepidotes.
Leufuichthys[35]	Gen. et sp.	Valid	Gallo Calvo Kellner	Turonian/Coniacian	Portezuelo Formation	Argentina	A clupeomorph.
Maorigadus ukrainicus[22]	Sp. nov	Valid	Schwarzhans Bratishko	Paleocene (Selandian)		Ukraine	A member of Gadiformes of uncertain phylogenetic placement, related to the family Muraenolepididae; a species of Maorigadus.
Mene iberica[19]	Sp. nov	Valid	Brzobohatý Nolf	Eocene (Bartonian)		Spain	A species of Mene.
"genus Merlucciidarum" antiquus[22]	Sp. nov	Valid	Schwarzhans Bratishko	Paleocene (Selandian)		Ukraine	A member of Merlucciidae of uncertain generic assignment.
Mimipiscis[36]	Nom. nov.	Valid	Choo	Upper Devonian	Gogo Formation	Australia	A new generic name for Mimia Gardiner & Bartram (1977). The type species is Mimipiscis toombsi (Gardiner & Bartram 1977).
Mimipiscis bartrami[36]	Species	Valid	Choo	Upper Devonian	Gogo Formation	Australia	A second species of Mimipiscis.
Neocybium parvidentatum[37]	Species	Valid	Monsch Bannikov	Eocene (Bartonian)		Kazakhstan	A Spanish mackerel, a species of Neocybium.
Ogilbia luzanensis[22]	Sp. nov	Valid	Schwarzhans Bratishko	Paleocene (Danian to Selandian)		Denmark  Greenland  Ukraine	A brotula, a species of Ogilbia.
Onuxodon yebraensis[19]	Sp. nov	Valid	Brzobohatý Nolf	Eocene (Bartonian)		Spain	A pearlfish, a species of Onuxodon.
Pardachirus izarrae[19]	Sp. nov	Valid	Brzobohatý Nolf	Eocene (Bartonian)		Spain	A sole, a species of Pardachirus.
Pavarottia[38]	Gen. et sp. nov	Valid	Bannikov & Zorzin	Eocene	Monte Bolca locality	Italy	A member of Percoidei of uncertain phylogenetic placement. The type species is Pavarottia lonardonii.
Pesciarhamphus[39]	Sp. nov	Valid	Tyler & Bannikov	Eocene (Ypresian or Lutetian)	Monte Bolca locality	Italy	An aulorhamphid gasterosteiform. The type species is Pesciarhamphus carnevalei.
Pietschellus[40]	Gen. et sp.	Valid	Bannikov Carnevale	Ypresian	Monte Bolca	Italy	A percomorph.
Robertannia[41]	Gen. et sp. nov	Valid	Bannikov	Eocene	Monte Bolca locality	Italy	A member of Percoidei of uncertain phylogenetic placement, related to Hendrixella grandei. The type species is Robertannia sorbiniorum.
Scheenstia[42]	Gen. et sp.	Valid	López-Arbarello Sferco	Late Jurassic		Germany	A semionotiform.
Sharfia[43]	Gen. et sp.	Valid	Pietsch Carnevale	Eocene (late Ypresian)	Monte Bolca locality	Italy	A goosefish. The type species is Sharfia mirabilis.
Sinosaurichthys[44]	Gen. et 3 sp.	Valid	Wu Sun et al.[CAL 6]	Anisian	Guanling Formation	China	A saurichthyid.
Sorbinichthys africanus[45]	Species	Valid	Murray Wilson	Cenomanian or Turonian	Akrabou Formation	Morocco	A species of Sorbinichthys.
"genus Sparidarum" spatiatus[22]	Sp. nov	Valid	Schwarzhans Bratishko	Paleocene (Selandian)		Ukraine	A member of Sparidae of uncertain generic assignment.
Spinofacia[27]	Gen. et sp.	Valid	Mickle	Carboniferous	Manning Canyon Shale Formation	USA	A ’palaeoniscoid’.
Steurbautichthys[46]	Gen. et comb. nov	Valid	Taverne	Middle Jurassic (Aalenian to Bathonian)	Stanleyville Formation, Songa Limestones	Democratic Republic of the Congo	A relative of Ankylophorus; a new genus for "Pholidophorus" aequatorialis de Saint-Seine & Casier (1962).
Tarkus[47]	Gen. et sp. nov	Valid	Carnevale Pietsch	Eocene (late Ypresian)	Monte Bolca locality	Italy	A member of Ogcocephalidae. The type species is Tarkus squirei.

Notes

1. Gai, Donoghue, Zhu, Janvier, Stampanoni.
2. Underwood, Goswami Prasad, Verma and Flynn
3. Fischer, Voigt, Schneider, Buchwitz and Voigt
4. Pradel, Tafforeau, Maisey, Janvier.
5. Downs, Daeschler, Jenkins and Shubin.
6. Wu, Sun, Xu, Hao, Jiang, Sun.

References

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13. Alexander Ivanov, Tiiu Märss and Anne Kleesment (2011). "A new elasmobranch Karksiodus mirus gen. et sp. nov. from the Burtnieki Regional Stage, Middle Devonian of Estonia". Estonian Journal of Earth Sciences 60 (1): 22–30. doi:10.3176/earth.2011.1.02.
14. Alan Pradel, Paul Tafforeau, John G. Maisey and Philippe Janvier (2011). "A New Paleozoic Symmoriiformes (Chondrichthyes) from the Late Carboniferous of Kansas (USA) and Cladistic Analysis of Early Chondrichthyans". PLoS ONE 6 (9): e24938. Bibcode:2011PLoSO...624938P. doi:10.1371/journal.pone.0024938.
15. Charlie J. Underwood and David J. Ward (2011). "New hemigaleid shark from the late Eocene of Wadi Al-Hitan, Egypt". Journal of Vertebrate Paleontology 31 (3): 707–711. doi:10.1080/02724634.2011.563763.
16. Shawn A. Hamm and David J. Cicimurri (2011). "Early Coniacian (late Cretaceous) selachian fauna from the basal Atco Formation, Lower Austin Group, northcentral Texas". Paludicola 8 (3): 107–127.
17. Susan Turner and Carole J. Burrow (2011). "A Lower Carboniferous xenacanthiform shark from Australia". Journal of Vertebrate Paleontology 31 (2): 241–257. doi:10.1080/02724634.2011.550359.
18. Claire Derycke and Daniel Goujet (2011). "Multicuspidate shark teeth associated with chondrichthyan and acanthodian scales from the Emsian (Devonian) of southern Algeria" (PDF). Geodiversitas 33 (2): 209–226. doi:10.5252/g2011n2a1.
19. Rostislav Brzobohatý and Dirk Nolf (2011). "Fish otoliths from the Middle Eocene (Bartonian) of Yebra de Basa, province of Huesca, Spain". Bulletin de l’Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre 81: 279–295.
20. Louis Taverne (2011). "Les poissons du Santonien (Crétacé supérieur) d’Apricena (Italie du Sud). 3°. Apricenaichthys italicus gen. et sp. nov. (Teleostei, Polymixiiformes)" (PDF). Bollettino del Museo Civico di Storia Naturale di Verona 35: 19–31.
21. Jesús Alvarado-Ortega and Paulo M. Brito (2011). "A new species of Araripichthys (Teleostei, Elopocephala) from the Tlayúa Formation (Cretaceous, Albian), Mexico". Journal of Vertebrate Paleontology 31 (6): 1376–1381. doi:10.1080/02724634.2011.602779.
22. Werner Schwarzhans and Andriy Bratishko (2011). "The otoliths from the middle Paleocene of Luzanivka (Cherkasy district, Ukraine)". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 261 (1): 83–110. doi:10.1127/0077-7749/2011/0154.
23. Giorgio Carnevale, Stephen J. Godfrey and Theodore W. Pietsch (2011). "Stargazer (Teleostei, Uranoscopidae) cranial remains from the Miocene Calvert Cliffs, Maryland, U.S.A. (St. Marys Formation, Chesapeake Group)". Journal of Vertebrate Paleontology 31 (6): 1200–1209. doi:10.1080/02724634.2011.606856.
24. James C. Tyler and Alexandre F. Bannikov (2011). "A new species of Aulorhamphus fish from the Eocene of Monte Bolca, Italy (†Aulorhamphidae, basal Gasterosteiformes)". Studi e ricerche sui giacimenti terziari di Bolca, XIII - Miscellanea paleontologica 10: 37–45.
25. Guang-Hui Xu, Ke-Qin Gao (2011). "A new scanilepiform from the Lower Triassic of northern Gansu Province, China, and phylogenetic relationships of non-teleostean Actinopterygii". Zoological Journal of the Linnean Society 161 (3): 595–612. doi:10.1111/j.1096-3642.2010.00645.x.
26. Sergio Bogan, Louis Taverne and Federico L. Agnolin (2011). "Description of a new aspidorhynchid fish, Belonostomus lamarquensis sp. nov. (Halecostomi, Aspidorhynchiformes), from the continental Upper Cretaceous of Patagonia, Argentina". Bulletin de l’Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre 81: 235–245.
27. Mickle, K.E. (2011). "The early actinopterygian fauna of the Manning Canyon Shale Formation (upper Mississippian, lower Pennsylvanian) of Utah, U.S.A.". Journal of Vertebrate Paleontology 31 (5): 962–980. doi:10.1080/02724634.2011.595622.
28. Alexandre F. Bannikov, Giorgio Carnevale and N. V. Parin (2011). "The new family Caucasichthyidae (Pisces, Perciformes) from the Eocene of the North Caucasus". Paleontological Journal 45 (1): 83–89. doi:10.1134/S0031030111010047.
29. Detlev Thies and Rolf Bernhard Hauff (2011). "A new species of Dapedium LEACH, 1822 (Actinopterygii, Neopterygii, Semionotiformes) from the Early Jurassic of South Germany" (PDF). Palaeodiversity 4: 185–221.
30. Victoria M. Arbour; Philip J. Currie (2011). "An istiodactylid pterosaur from the Upper Cretaceous Nanaimo Group, Hornby Island, British Columbia, Canada". Canadian Journal of Earth Sciences 48 (1): 63–69. Bibcode:2011CaJES..48...63S. doi:10.1139/E10-083.
31. Romain Vullo, Eric Buffetaut and Michael J. Everhart (2012). "Reappraisal of Gwawinapterus beardi from the Late Cretaceous of Canada: a saurodontid fish, not a pterosaur". Journal of Vertebrate Paleontology 32 (5): 1198–1201. doi:10.1080/02724634.2012.681078.
32. Jason P. Downs, Edward B. Daeschler, Farish A. Jenkins Jr. and Neil H. Shubin (2011). "A new species of Laccognathus (Sarcopterygii, Porolepiformes) from the Late Devonian of Ellesmere Island, Nunavut, Canada". Journal of Vertebrate Paleontology 31 (5): 981–996. doi:10.1080/02724634.2011.599462.
33. Louis Taverne (2011). "Les poissons crétacés de Nardò. 33°. Lecceclupea ehiravaensis gen. et sp. nov. (Teleostei, Clupeidae)" (PDF). Bollettino del Museo Civico di Storia Naturale di Verona 35: 3–17.
34. Peter L. Forey, Adriana López-Arbarello, and Norman MacLeod (2011). "A new species of Lepidotes (Actinopterygii: Semiontiformes) from the Cenomanian (Upper Cretaceous) of Morocco". Palaeontologia Electronica 4 (1): 7A:1–12.
35. Valéria Gallo, Jorge O. Calvo and Alexander W.A. Kellner (2011). "First record of a clupeomorph fish in the Neuquén Group (Portezuelo Formation), Upper Cretaceous of Patagonia, Argentina". Cretaceous Research 32 (2): 223–235. doi:10.1016/j.cretres.2010.12.006.
36. Brian Choo (2011). "Revision of the actinopterygian genus Mimipiscis (=Mimia) from the Upper Devonian Gogo Formation of Western Australia and the interrelationships of the early Actinopterygii". Earth and Environmental Science Transactions of the Royal Society of Edinburgh 102 (2): 77–104. doi:10.1017/S1755691011011029.
37. Kenneth A. Monsch and Alexandre F. Bannikov (2011). "New taxonomic synopses and revision of the scombroid fishes (Scombroidei, Perciformes), including billfishes, from the Cenozoic of territories of the former USSR". Earth and Environmental Science Transactions of the Royal Society of Edinburgh 102 (4): 253–300. doi:10.1017/S1755691011010085.
38. Alexandre F. Bannikov and Roberto Zorzin (2011). "A new priacanthid-like percoid fish (Perciformes) from the Eocene of Bolca, Italy". Studi e ricerche sui giacimenti terziari di Bolca, XIII - Miscellanea paleontologica 10: 17–27.
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