Pomatiidae

Pomatiidae
An apertural view of a shell and operculum of Tudorella sulcata
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
(unranked):	clade Caenogastropoda clade Littorinimorpha
Superfamily:	Littorinoidea
Family:	Pomatiidae Newton, 1891 (1828)
Synonyms
Pomatiasidae

The family Pomatiidae is a taxonomic family of small operculate land snails, terrestrial gastropod mollusks that can be found over the warmer parts of the Old World. In the older literature, this family is designated as Pomatiasidae.

This family is a lineage closely related to the Littorinidae (periwinkles) common in coastal habitat. They have adapted to terrestrial life and are thus sometimes called "land winkles".^[1]

They are defined by a chalky operculum at the rear end of the body, the shape of their thick shell and their mouth and a characteristic spiral sculpture. The sexes are separate and can sometimes be recognised because the female shell is slightly larger than the male shell.

Taxonomy

Tropidophora fimbriata haemostoma.

According to the Taxonomy of the Gastropoda (Bouchet & Rocroi, 2005), the family Pomatiidae consists of two subfamilies:^[2]

• subfamily Pomatiinae Newton, 1891 (1828) - synonyms: Cyclostomatidae Menke, 1828; Cyclotopsinae Kobelt & Möllendorff, 1898; Ericiidae Wenz, 1915
• subfamily Annulariinae Henderson & Bartsch, 1920^[3]
  • tribe Annulariini Henderson & Bartsch, 1920 - synonyms: Licininae Gray, 1857; Chondropomatinae Henderson & Bartsch, 1920
  • tribe Adamsiellini Henderson & Bartsch, 1920
  • tribe Choanopomatini Thiele, 1929
  • tribe Cistulopsini H. B. Baker, 1924 - synonym: Cistulinae L. Pfeiffer, 1858
  • tribe Rhytidopomatini Henderson & Bartsch, 1920

Genera

Genera within the family Pomatiidae include:

subfamily Pomatiinae

• Cyclostoma Lamarck, 1899^[2]
• Cyclotopsis Blanford, 1864^[2]
• Ericia Partiot, 1848^[2]
• Georgia Bourguignat, 1882^[4]
• Guillainia Crosse, 1884^[5]
• Leonia Gray, 1850^[6]
• Lithidion Gray, 1850^[7]
• Otopoma Gray, 1850^[8]
• Pomatias Studer, 1789 - type genus of the family Pomatiidae^[2]
• Tropidophora Troschel, 1847^[9]
• Tudorella Fischer, 1885^[10]

subfamily Annulariinae

(to be sorted to tribes)

• Abbottella Henderson & Bartsch, 1920^[11]
• Articulipoma Bartsch, 1946^[11]
• Chondropomium Henderson & Bartsch, 1920^[11]
• Parachondria Dall, 1905^[11] - originally described as a subgenus^[12]

tribe Annulariini

• Annularia Schumacher, 1817 - type genus of the subfamily Annulariinae^[2]
• Chondropoma L. Pfeiffer, 1847^[2][13]
• Licina Gray, 1847^[2]

tribe Adamsiellini

• Adamsiella L. Pfeiffer, 1851 - type genus of the tribe Adamsiellini^[2]

tribe Choanopomatini

• Choanopoma L. Pfeiffer, 1847 - type genus of the tribe Choanopomatini^[2]

tribe Cistulopsini

• Cistula Gray, 1850^[2]
• Cistulops H. B. Baker, 1924 - type genus of the tribe Cistulopsini^[2]

tribe Rhytidopomatini

• Rhytidopoma Sykes, 1901 - type genus of the tribe Rhytidopomatini^[2][13]

to be sorted to subfamilies:

• Annularisca Henderson & Bartsch, 1920^[13]
• Annularita Henderson & Bartsch, 1920^[13]
• Annularodes Henderson & Bartsch, 1920^[13]
• Annularops Henderson & Bartsch, 1920^[13]
• Bermudezia Torre & Bartsch, 1941^[13]
• Blaesospira Crosse, 1890^[13]
• Chondropomatus Henderson & Bartsch, 1920^[13]
• Chondropometes Henderson & Bartsch, 1920^[13]
• Chondrothyra Henderson & Bartsch, 1920^[13]
• Chondrothyrella Torre & Bartsch, 1938^[13]
• Chondrothyrium Henderson & Bartsch, 1920^[13]
• Cubadamsiella Torre & Bartsch, 1941^[13]
• Dallsiphona Torre & Bartsch, 1941^[13]
• Diploma^[13] - an orth. error for Diplopoma? (original research)
  • Diploma arangoi (Jaume, 1984)^[13]
  • Diploma architectonica (Gundlach in Pfeiffer, 1859)^[13]
  • Diploma songoensis (Torre & Bartsch, 1941)^[13]
  • Diploma torrei (Ramsden, 1915)^[13]
  • Diploma varonai (Jaume, 1984)^[13]
  • Diploma zayasi (Jaume, 1984)^[13]
• Diplopoma L. Pfeiffer, 1859^[14]
• Eutudora Henderson & Bartsch, 1920^[13]
• Eutudorops Henderson & Bartsch, 1920^[13]
• Guajaibona Torre & Bartsch, 1941^[13]
• Gundlachtudora Torre & Bartsch, 1941^[13]
• Hendersonina Torre & Bartsch, 1938^[13]
• Jaumeia Aguayo, 1943^[13]
• Juannularia Torre & Bartsch, 1941^[13]
• Limadora Torre & Bartsch, 1941^[13]
• Limadorex Torre & Bartsch, 1941^[13]
• Meganipha Thompson, 1978
• Opisthocoelicum Torre & Bartsch, 1941^[13]
• Opisthosiphon Dall, 1905^[12][13]
• Ramsdenia Preston, 1913^[13]
• Rhytidothyra Henderson & Bartsch, 1920^[13]
• Subannularia Torre & Bartsch, 1941^[13]
• Torrella Henderson & Bartsch, 1920^[13]
• Torrilla^[13] (spelling error?)
  • Torrilla emmae Jaume & Sánchez de Fuentes, 1943^[13]
  • Torrilla torreiana (Gundlach in Arango, 1878)^[13]
  • Torrilla trinidadensis Torre & Bartsch, 1941^[13]
• Troschelvindex Baker, 1924^[13]
• Tudora Baird, 1850
• Tudorina Torre & Bartsch, 1941^[13]
• Wrightudora Torre & Bartsch, 1941^[13]
• Xenopoma Crosse, 1890^[13]
• Xenopoides^[13]
  • Xenopoides delicatulum Torre & Bartsch, 1941^[13]

References

1. Journal of Molluscan Studies, GENETIC AND MORPHOLOGICAL VARIATION IN THE LAND WINKLE POMATIAS ELEGANS (MÜLLER) (CAENOGASTROPODA: POMATIASIDAE) Volume 67, Issue 2, pp 145–152 (http://mollus.oxfordjournals.org/content/67/2/145.abstract) Accessed 2014-9-2
2. Bouchet P., Rocroi J.-P., Frýda J., Hausdorf B., Ponder W., Valdés Á. & Warén A. (2005). "Classification and nomenclator of gastropod families". Malacologia: International Journal of Malacology (Hackenheim, Germany: ConchBooks) 47 (1-2): 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
3. Henderson J. B. Jr. & Bartsch P. (1920). "A classification of the American operculate land mollusks of the family Annulariidae". Proceedings of the United States National Museum 58: 49-82. page 54.
4. Georgia in GBIF
5. Guillainia in Bionames.org, accessed 12 december 2015.
6. "Genus summary for Leonia". AnimalBase, accessed 28 march 2011.
7. Lithidion in GBIF
8. Otopoma Gray 1850 - a few words to add to a 150 years old debate (Mollusca: Caenogastropoda: Cyclophoridae)
9. KENNETH C. EMBERTON Cryptic, genetically extremely divergent, polytypic, convergent, and polymorphic taxa in Madagascan Tropidophora (Gastropoda: Pomatiasidae) Biological Journal of the Linnean Society - Volume 55, Issue 3, pages 183–208, July 1995
10. "Genus summary for Tudorella". AnimalBase, accessed 28 march 2011.
11. Watters G. T. (2012). "Hispaniolan Annulariidae (Gastropoda), primarily from the Barahona Peninsula: new taxa and notes". The Nautilus 126: 1-14.
12. Dall W. H. (1905). "An arrangement of the American Cyclostomatidae, with a revision of the nomenclature". Proceedings of the Malacological Society of London 6: 208-210.
13. "Mollusca". Diversidad Biológica Cubana, accessed 23 March 2011.
14. Robinson D. G., Hovestadt A., Fields A. & Breure A. S. H. (July 2009). "The land Mollusca of Dominica (Lesser Antilles), with notes on some enigmatic or rare species". Zoologische Mededelingen 83 http://www.zoologischemededelingen.nl/83/nr03/a13

Further reading

• Bartsch P. (1946). "The operculate land mollusks of the family Annulariidae of the Island of Hispaniola and the Bahama Archipelago". United States National Museum Bulletin No. 192, 264 pp., 38 pls.
• Pfenninger, M.; Véla, E.; Jesse, R.; Elejalde, M. A.; Liberto, F.; Magnin, F. D. R.; Martínez-Ortí, A. (2010). "Temporal speciation pattern in the western Mediterranean genus Tudorella P. Fischer, 1885 (Gastropoda, Pomatiidae) supports the Tyrrhenian vicariance hypothesis". Molecular Phylogenetics and Evolution 54 (2): 427–436. doi:10.1016/j.ympev.2009.09.024. PMID 19772926.
• Watters G. T. (2006). The Caribbean Land Snail Family Annulariidae: A revision of the higher taxa and a catalog of the species. 1-557, figs 1-10, map 1-56, Appendix A: 1-3. Backhuys Publishers, Leiden.

External links

• "Pomatiasidae". National Center for Biotechnology Information (NCBI).