Baltic amber

Different colours of Baltic amber.

Typical beach sand on the Baltic Sea where amber is often washed up.

Fishing for amber at the seacoast, Mikoszewo, close to Gdańsk, Poland.

The Baltic region is home to the largest known deposit of amber, called Baltic amber or succinite. It dates from 44 million years ago (during the Eocene epoch).^[1] It has been estimated that these forests created more than 100,000 tons of amber.^[2] Today, more than 90% of the world's amber comes from Kaliningrad Oblast of Russia. It's a major source of income for the region; the local Kaliningrad Amber Combine extracted 250 tonnes of it in 2014,^[3] 400 tonnes in 2015.^[4]

The term "Baltic amber" is generic, so amber from the Bitterfeld brown coal mines in Saxony (Eastern Germany) goes under the same name. Bitterfeld amber was previously believed to be only 20–22 million years old (Miocene), but a comparison of the animal inclusions revealed that it is most probably genuine Baltic amber that has only been redeposited in a Miocene deposit.^[5] Other sources of Baltic amber have been listed as coming from Poland and Russia.

Because Baltic amber contains about 8% succinic acid, it is also termed succinite.

It was thought since the 1850s that the resin that became amber was produced by the tree Pinites succinifer, but research in the 1980s came to the conclusion that the resin originates from several species. More recently, it has been proposed, on the evidence of Fourier-transform infrared microspectroscopy (FTIR) analysis of amber and resin from living trees, that conifers of the family Sciadopityaceae were responsible.^[2] The only extant representative of this family is the Japanese umbrella pine, Sciadopitys verticillata.

Numerous extinct genera and species of plants and animals have been discovered and scientifically described from inclusions in Baltic amber.^[6] Baltic amber includes the most species-rich fossil insect fauna discovered to date.

Paleobiology

Numerous organisms have been described from amber specimens including:

Fossil brentid beetle

Flora

• Notoscyphus balticus^[7]
• Rhizomnium dentatum Heinrichs et al, 2014^[8]

Fauna

Agroecomyrmex Wheeler, 1910[9] Aphaenogaster mersa Wheeler, 1915[9] Aphaenogaster oligocenica Wheeler, 1915[9] Aphaenogaster sommerfeldti (Mayr, 1868)[9] Arostropsis Yunakov & Kirejtshuk, 2011[10] Aspidopleura Gibson, 2009[11] Asymphylomyrmex Wheeler, 1915[9] Baltimartyria Skalski, 1995 Baltocteniza Eskov & Zonstein, 2000[12] Brevivulva Gibson, 2009[11] Deinodryinus areolatus (Ponomarenko, 1975)[13] Deinodryinus velteni Guglielmino & Olmi, 2011[13] Diochus electrus Chatzimanolis & Engel, 2011[14] Electrinocellia (Carpenter) Engel, 1995[15] Electrocteniza Eskov & Zonstein, 2000[12]

Electropodagrion Azar & Nel, 2008[16] Electrostephanus Brues, 1933[17] Elektrithone Makarkin, Wedmann, & Weiterschan, 2014[18] Epiborkhausenites Skalski, 1973[19] Glisachaemus Szwedo, 2007[20] Gracillariites Kozlov, 1987 Metanephrocerus collini Carpenter & Hull, 1939[21] Metanephrocerus groehni Kehlmaier & Skevington, 2014[21] Metanephrocerus hoffeinsorum Kehlmaier & Skevington, 2014[21] Electrocrania Kuznezov, 1941 Fibla carpenteri Engel, 1995[15] Metapelma archetypon Gibson, 2009[11] Micropterix gertraudae Kurz & Kurz, 2010 Neanaperiallus Gibson, 2009[11] Palaeovespa baltica Cockerell, 1909[22]

Palaeovespa socialis Pionar, 2005[23] Prolyonetia Kusnetzov, 1941 Propupa Stworzewicz & Pokryszko, 2006[24] Pseudogarypus synchrotron Henderickx, 2012[25] Stigmellites baltica (Kozlov, 1988) (Lepidopteran leaf mines) Xylolaemus sakhnovi Alekseev & Lord, 2014[26] Succinipatopsis Poinar, 2000[27] Yantaromyrmex constricta (Mayr, 1868)[28] Yantaromyrmex geinitzi (Mayr, 1868)[28] Yantaromyrmex samlandica (Wheeler, 1915)[28]

See also

• Amber Coast
• Dominican amber

References

1. Ritzkowski, S. (1997). "K-Ar-Altersbestimmungen der Bernsteinführenden Sedimente des Samlandes (Paläogen, Bezirk Kaliningrad)". Metalla (Bochum) 66 (Sonderheft): 19–23.
2. Wolfe, A. P.; Tappert, R.; Muehlenbachs, K.; Boudreau, M.; McKellar, R. C.; Basinger, J. F.; Garrett, A. (2009). "A New Proposal Concerning the Botanical Origin of Baltic Amber". Proceedings of the Royal Society B 276 (1672): 3403–3412. doi:10.1098/rspb.2009.0806. PMC 2817186. PMID 19570786.
3. "Russia: Amber covers beach after storm". BBC. 9 January 2015.
4. "Amber mining". Amber Museum Kaliningrad.
5. Dunlop, J. A.; Giribet, G. (2003). "The first Fossil Cyphophthalmid (Arachnida: Opiliones), from Bitterfeld Amber, Germany". Journal of Arachnology 31 (3): 371–378. doi:10.1636/h03-03.
6. Weitschat, W.; Wichard, W. (2002). Atlas of Plants and Animals in Baltic Amber. Pfeil. ISBN 978-3931516949.
7. Heinrichs, J; Schmidt, AR; Schäfer-Verwimp, A; Gröhn, C; Renner, MAM (2015). "The leafy liverwort Notoscyphus balticus sp. nov. (Jungermanniales) in Eocene Baltic amber". Review of Palaeobotany and Palynology 217: 39–44. doi:10.1016/j.revpalbo.2015.02.006.
8. Heinrichs, J; Hedenäs, L; Schäfer-Verwimp, A; Feldberg, K; Schmidt, AR (2014). "An in situ preserved moss community in Eocene Baltic amber". Review of Palaeobotany and Palynology 210: 113–118. doi:10.1016/j.revpalbo.2014.08.005.
9. Wheeler, W. M. (1914). "The Ants of the Baltic Amber". Schriften der Physikalisch-Ökonomischen Gesellschaft zu Königsberg 55 (4): 56–59.
10. Yunakov, N.N.; Kirejtshuk, A.G. (2011). "New genus and species of broad-nosed weevils from Baltic amber and notes on fossils of the subfamily Entiminae (Coleoptera, Curculionidae)". ZooKeys 160: 73–96. doi:10.3897/zookeys.160.2108. PMC 3253632. PMID 22303121.
11. Gibson, G. A. P. (2009). "Description of three New Genera and four New Species of Neanastatinae (Hymenoptera, Eupelmidae) from Baltic Amber, with Discussion of their Relationships to Extant Taxa". ZooKeys 20: 175–214. doi:10.3897/zookeys.20.161.
12. Eskov, K. Y.; Zonstein, S. L. (2000). "The First Ctenizoid Mygalomorph Spiders from Eocene Baltic Amber (Araneida: Mygalomorphae: Ctenizidae)". Paleontological Journal 34 (3): S268–S274. Part 1; part 2, PDF.
13. Guglielmino, A.; Olmi, M. (2011). "Revision of fossil species of Deinodryinus, with description of a new species (Hymenoptera, Dryinidae)". ZooKeys 130: 495–504. doi:10.3897/zookeys.130.1326. PMC 3260777. PMID 22259295.
14. Chatzimanolis, S.; Engel, M. S. (2011). "A new species of Diochus from Baltic amber (Coleoptera, Staphylinidae, Diochini)" (PDF). ZooKeys 138: 65–73. doi:10.3897/zookeys.138.1896.
15. Engel, M. S. (1995). "A New Fossil Snake-Fly Species from Baltic Amber (Raphidioptera: Inocelliidae)". Psyche 102 (3–4): 187–193. doi:10.1155/1995/23626.
16. Azar, D.; Nel, A. (2008). "First Baltic amber megapodagrionid damselfly (Odonata: Zygoptera)". Annales de la Société Entomologique de France. New Series 44 (4): 451–457. doi:10.1080/00379271.2008.10697580. – via Taylor & Francis (subscription required)
17. Engel, M.S.; Ortega-Blanco, J. (2008). "The fossil crown wasp Electrostephanus petiolatus Brues in Baltic Amber (Hymenoptera, Stephanidae): designation of a neotype, revised classification, and a key to amber Stephanidae". ZooKeys 4: 55–64. doi:10.3897/zookeys.4.49.
18. Makarkin, VN; Wedmann, S; Weiterschan, T (2014). "First record of the family Ithonidae (Neuroptera) from Baltic amber". Zootaxa 3796 (2): 385–393. doi:10.11646/zootaxa.3796.2.10.
19. Skalski, A. W. (1973). "Studies on the Lepidoptera from Fossil Resins. Part II. Epiborkhausenites obscurotrimaculatus gen. et sp. nov. (Oecophoridae) and a Tineid-Moth Discovered in the Baltic Amber" (PDF). Acta Palaeontologica Polonica 18 (1): 153–160.
20. Szwedo, J. (2007). "Glisachaemus jonasdamzeni gen. et sp. nov. of Cixiidae from the Eocene Baltic amber (Hemiptera: Fulgoromorpha)" (PDF). Alavesia 1: 109–116.
21. Kehlmaier, C; Dierick, M; Skevington, JH (2014). "Micro-CT studies of amber inclusions reveal internal genitalic features of big-headed flies, enabling a systematic placement of Metanephrocerus Aczel, 1948 (Insecta: Diptera: Pipunculidae)". Arthropod Systematics & Phylogeny 72 (1): 23–36.
22. Cockerell, T. D. A. (1906). "Fossil Hymenoptera from Florissant, Colorado" (PDF). Bulletin of the Museum of Comparative Zoology 50 (2).
23. Poinar, G. (2005). "Fossil Trigonalidae and Vespidae (Hymenoptera) in Baltic Amber". Proceedings of the Entomological Society of Washington 107 (1): 55–63.
24. Stworzewicz, E.; Pokryszko, B. M. (2006). "Eocene Terrestrial Snails (Gastropoda) from Baltic Amber" (PDF). Annales Zoologici 56 (1): 215–224.
25. Henderickx, H.; Tafforeau, P.; Soriano, C (2012). "Phase-contrast synchrotron microtomography reveals the morphology of a partially visible new Pseudogarypus in Baltic amber (Pseudoscorpiones: Pseudogarypidae)". Palaeontologia Electronica 15 (2;17A,11p): 1–11.
26. Alekseev, VI; Lord, NP (2014). "A new species of Xylolaemus (Coleoptera: Zopheridae: Colydiinae) from Baltic amber". Baltic Journal of Coleopterology 14 (1): 97–102. ISSN 1407-8619.
27. Poinar, G. Jr. (2000). "Fossil Onychophorans from Dominican and Baltic Amber: Tertiapatus dominicanus n.g., n.sp. (Tertiapatidae n.fam.) and Succinipatopsis balticus n.g., n.sp. (Succinipatopsidae n.fam.) with a Proposed Classification of the Subphylum Onychophora". Invertebrate Biology 119 (1): 104–9. doi:10.1111/j.1744-7410.2000.tb00178.x.
28. Dlussky, GM; Dubovikoff, DA (2013). PDF "Yantaromyrmex gen. n. – a new ant genus (Hymenoptera Formicidae) from Late Eocene ambers of Europe" Check |url= value (help) (PDF). Caucasian Entomological Bulletin 9: 305–314.