Cadulus

Cadulus
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Scaphopoda
Order:	Gadilida
Family:	Gadilidae
Genus:	Cadulus Philippi, 1844 [1]
Species
51, see text.

Cadulus is a large genus of small tusk shells in the family Gadilidae. It contains over fifty-one described species.^[2]

Species

• Cadulus acus Dall, 1889
• Cadulus aequalis Dall, 1881
• Cadulus agassizii Dall, 1881
• Cadulus albicomatus Dall, 1890
• Cadulus amiantus Dall, 1889
• Cadulus amphora Jeffreys, 1883
• Cadulus ampullaceus Watson, 1879
• Cadulus aratus Hedley, 1899
• Cadulus arctus Henderson, 1920
• Cadulus artatus Locard, 1897
• Cadulus atlanticus Henderson, 1920
• Cadulus attenuatus Monterosato, 1875
• Cadulus austinclarki Emerson, 1951
• Cadulus californicus (Pilsbry and Sharp, 1898)
• Cadulus campylus Melvill, 1906
• Cadulus catharus Henderson, 1920
• Cadulus chuni Jaeckel, 1932
• Cadulus colliverae Lamprell & Healy, 1998
• Cadulus colubridens Watson, 1879
• Cadulus congruens Watson, 1879
• Cadulus cucurbitus Dall, 1881
• Cadulus curtus Watson, 1879
• Cadulus cyathoides Jaeckel, 1932
• Cadulus cylindratus Jeffereys, 1877
• Cadulus delicatulus Suter, 1913
• Cadulus deschampsi Scarabino, 2008
• Cadulus deverdensis Scarabino, 2008
• Cadulus eliezeri Caetano, Scarabino & Absalão, 2006
• Cadulus euloides Melvill & Standen, 1901
• Cadulus exiguus Watson, 1879
• Cadulus florenciae Scarabino, 1995
• Cadulus foweyensis Henderson, 1920
• Cadulus gibbus Jeffreys, 1883
• Cadulus glans Scarabino, 1995
• Cadulus gracilis Jeffreys, 1877
• Cadulus greenlawi Dall, 1889
• Cadulus iota Henderson, 1920
• Cadulus jeffreysi (Monterosato, 1875)
• Cadulus labeyriei Scarabino, 1995
• Cadulus lemniscoides Scarabino, 2008
• Cadulus loyaltyensis Scarabino, 2008
• Cadulus lunulus Dall, 1881
• Cadulus macleani Emerson, 1978
• Cadulus mayori (J. B. Henderson, 1920)
• Cadulus miamiensis J. B. Henderson, 1920
• Cadulus minusculus Dall, 1889
• Cadulus monterosatoi Locard, 1897
• Cadulus nerta Caetano, Scarabino & Absalão, 2006
• Cadulus obesus Watson, 1879
• Cadulus occiduus Verco, 1911
• Cadulus ovulum (Philippi, 1844)
• Cadulus parvus J. B. Henderson, 1920
• Cadulus platei Jaeckel, 1932
• Cadulus platensis Henderson, 1920
• Cadulus podagrinus Henderson, 1920
• Cadulus propinquus Sars G.O., 1878
• Cadulus providensis Henderson, 1920
• Cadulus quadridentatus Dall, 1881
• Cadulus quadrifissatus (Pilsbry and Sharp, 1898)
• Cadulus rastridens Watson, 1879
• Cadulus regularis Henderson, 1920
• Cadulus rocroii Scarabino, 2008
• Cadulus rossoi Nicklès, 1979
• Cadulus rudmani Lamprell & Healy, 1998
• Cadulus scarabinoi Steiner & Kabat, 2004
• Cadulus siberutensis Jaeckel, 1932
• Cadulus simillimus Watson, 1879
• Cadulus sofiae Scarabino, 1995
• Cadulus stearnsii (Pilsbry and Sharp, 1898)
• Cadulus subfusiformis (M. Sars, 1865)
• Cadulus teliger Finlay, 1927
• Cadulus tersus Henderson, 1920
• Cadulus tetraschistus (Watson, 1879)
• Cadulus tetrodon Pilsbry and Sharp, 1898
• Cadulus thielei Plate, 1908
• Cadulus tolmiei (Dall, 1897)
• Cadulus transitorius (J. B. Henderson, 1920) - inflated toothshell
• Cadulus tumidosus Jeffreys, 1877
• Cadulus unilobatus V. Scarabino & F. Scarabino, 2011
• Cadulus valdiviae Jaeckel, 1932
• Cadulus vincentianus Cotton & Godfrey, 1940
• Cadulus watsoni Dall, 1881
• Cadulus woodhousae Lamprell & Healy, 1998

Species brought into synonymy

• Cadulus aberrans Whiteaves, 1887 - aberrant toothshell: synonym of Gadila aberrans (Whiteaves, 1887)
• Cadulus booceras Tomlin, 1926: synonym of Siphonodentalium booceras (Tomlin, 1926)
• Cadulus carolinensis Bush, 1885: synonym of Polyschides carolinensis (Bush, 1885)
• Cadulus domingensis (d'Orbigny in Mörch, 1863): synonym of Gadila dominguensis (d’Orbigny, 1853)
• Cadulus carolinensis Bush, 1885: synonym of Polyschides carolinensis (Bush, 1885)
• Cadulus elongatus J. B. Henderson, 1920: synonym of Gadila elongata (Henderson, 1920)
• Cadulus fusiformis Pilsbry and Sharp, 1898: synonym of Gadila aberrans (Whiteaves, 1887)
• Cadulus grandis A. E. Verrill, 1884: synonym of Polyschides grandis (Verrill, 1884)
• Cadulus halius Henderson, 1920: synonym of Cadulus podagrinus Henderson, 1920
• Cadulus hepburni (Dall, 1897): synonym of Gadila aberrans (Whiteaves, 1887)
• Cadulus laevis (Brazier, 1877): synonym of Ditrupa gracillima Grube, 1878
• Cadulus martini Scarabino, 1995: synonym of Cadulus scarabinoi Steiner & Kabat, 2004
• Cadulus nitentior Arnold, 1903 †: synonym of Gadila aberrans (Whiteaves, 1887)
• Cadulus nitidus: synonym of Polyschides nitidus (Henderson, 1920)
• Cadulus pandionis (A. E. Verrill and S. Smith, 1880): synonym of Gadila pandionis (Verrill & Smith, 1880)
• Cadulus perpusillus (Sowerby, 1832): synonym of Gadila perpusilla (Sowerby in Broderip & Sowerby, 1832)
• Cadulus peruvianus Dall, 1908: synonym of Gadila peruviana (Dall, 1908)
• Cadulus poculum Dall, 1889: synonym of Gadila pocula (Dall, 1889)
• Cadulus rushii Pilsbry and Sharp, 1898: synonym of Polyschides rushii (Pilsbry & Sharp, 1898)
• Cadulus simpsoni Henderson, 1920: synonym of Gadila simpsoni Henderson, 1920
• Cadulus spectabilis A. E. Verrill, 1885: synonym of Polyschides spectabilis (A. E. Verrill, 1885)
• Cadulus strangulatus Locard, 1897: synonym of Gadila strangulata (Locard, 1897)
• Cadulus striatus Pilsbry and Sharp, 1898: synonym of Gadila striata (Pilsbry & Sharp, 1898)
• Cadulus subtilis Plate, 1908: synonym of Ditrupa gracillima Grube, 1878
• Cadulus verrilli Henderson, 1920: synonym of Gadila verrilli (Henderson, 1920)
• Cadulus viperidens Melvill & Standen, 1896: synonym of Dischides viperidens (Melvill & Standen, 1896)

References

1. Philippi R. A. (1844). Enumeratio molluscorum Siciliae cum viventium tum in tellure tertiaria fossilium, quae in itinere suo observavit. Vol. 2. Eduard Anton, Halle [Halis Saxorum] : iv + 303 p., pl. 13-28
2. Scarabino, V.; Gofas, S. (2012). Cadulus. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=138024 on 2012-07-16

• Powell A W B, New Zealand Mollusca, William Collins Publishers Ltd, Auckland, New Zealand 1979 ISBN 0-00-216906-1
• "Cadulus". Integrated Taxonomic Information System.
• Steiner G. & Kabat A.R. (2001). Catalogue of supraspecific taxa of Scaphopoda (Mollusca). Zoosystema 23 (3): 433-460