Enidae

Enidae
A live individual of Ena montana
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
(unranked):	clade Heterobranchia clade Euthyneura clade Panpulmonata clade Eupulmonata clade Stylommatophora informal group Orthurethra
Superfamily:	Enoidea
Family:	Enidae B. B. Woodward, 1903 (1880)
Genera
See text

Enidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks.

Anatomy

In this family, the number of haploid chromosomes lies between 21 and 25 (according to the values in this table).^[1]

Taxonomy

1998 taxonomy

The following subfamilies of Enidae are based on work by Schileyko (1998)^[2] and they are in use mainly by Russian malacologists:^[3]

• subfamily Pseudonapaeinae
• subfamily Chondrulopsininae
• subfamily Merdigerinae
• subfamily Buliminusinae
• subfamily Andronakiinae
• subfamily Retowskiinae
• subfamily Eninae - including Chondrula
• subfamily Multidentulini

2005 taxonomy

The following two subfamilies have been recognized in the taxonomy of Bouchet & Rocroi (2005):

• subfamily Eninae B. B. Woodward, 1903 (1880)
  • tribus Enini B. B. Woodward, 1903 (1880) - synonyms: Napaeinae A. J. Wagner, 1928; Jaminiinae Thiele, 1931; Pseudonapaeinae Schileyko, 1978; Retowskiinae Schileyko, 1978; Andronakiinae Schileyko, 1998
  • tribus Chondrulini Wenz, 1923
  • tribus Multidentulini Schileyko, 1978 - synonyms: Chondrulopsininae Schileyko, 1978; Merdigerinae Schileyko, 1984; Euchondrinae Schileyko, 1998
• subfamily Buliminusinae Kobelt, 1880 - synonym Buliminidae Pfeiffer, 1879

Genera

Genera within the family Enidae include:

subfamilia Eninae

• Boninena Habe, 1956

either in Enini or in Chondrulini

• Brephulopsis Lindholm, 1925^[3]
• Caucasicola Hesse, 1917^[3]
• Chondrus Cuvier, 1817^[3]
• Georginapaeus Schileyko, 1988^[3]
• Peristoma Krynicki, 1833^[3]
• Ramusculus Lindholm, 1925^[3]
• Thoanteus Lindholm, 1925^[3]
• tribus Enini
  • Akramowskiella Schileyko, 1984^[3]
  • Andronakia Lindholm, 1914^[3]
  • Clausilioides Lindholm, 1925^[3]
  • Ayna Páll-Gergely, 2009^[4]
  • Differena Schileyko, 1984^[3]
  • Ena Turton, 1831^[3] - type genus of the family Enidae
  • Geminula Lindholm, 1925^[3]
  • Imparietula Lindholm, 1925^[3]
  • Jaminia Risso, 1826
  • Laevozebrinus Lindholm, 1925^[3]
  • Ljudmilena Schileyko, 1984^[3]
  • Mastoides Westerlund, 1896^[3][5]
  • Megalena Hausdorf, 1999^[6] There is only one species Megalena crassa (Retowski, 1887) in the genus Megalena.
  • Napaeus Albers, 1850
  • Ottorosenia Muratov, 1992^[3]
  • Pseudochondrula Hesse, 1933^[3]
  • Pseudonapaeus Westerlund, 1887^[3]
  • Retowskia O. Boettger, 1881^[3]
  • Subzebrinus Westerlund, 1887^[3]
  • Turanena Lindholm, 1922^[3]
• tribus Chondrulini
  • Chondrula Beck, 1837^[3] - type genus of the tribe Chondrulini
  • Mastus Beck, 1837
  • Zebrina Held, 1837^[3]
• tribus Multidentulini
  • Chondrulopsina Lindholm, 1925^[3]
  • Euchondrus Boettger, 1883^[3]
  • Improvisa Schileyko, 1978^[3]
  • Multidentula Lindholm, 1925
  • Merdigera Held, 1837^[3]
  • Pentadentula Suvorov, 2006^[3]
  • Senaridenta Schileyko, 1978^[3]
  • Siraphoroides Schileyko, 1977^[3]

subfamily Buliminusinae

• Adzharia Hesse, 1933^[3]
• Buliminus Beck, 1837^[3] - type genus of the subfamily Buliminusinae
• Cyrenaeus Heller, 1971
• Mordania Bank & Neubert, 1998
• Paramastus Hesse, 1933
• Pene Pallary, 1929

subfamily ?

• † Balearena Altaba, 2007^[7] - type species † Balearena gymnesica Altaba, 2007^[7]
• Luchuena Habe, 1956

References

1. Barker G. M.: Gastropods on Land: Phylogeny, Diversity and Adaptive Morphology. in Barker G. M. (ed.): The biology of terrestrial molluscs. CABI Publishing, Oxon, UK, 2001, ISBN 0-85199-318-4. 1-146, cited pages: 139 and 142.
2. Schileyko A. A. (1998). Treatise on Recent terrestrial pulmonate molluscs. Part 2. Gastrocoptidea, Hypselostomatidae, Vertiginidae, Truncatellinidae, Pachynodidae, Enidae, Sagdidae. Ruthenica Supplement 2(2): 129-261.
3. Kantor Yu I., Vinarski M. V., Schileyko A. A. & Sysoev A. V.(published online on December 22, 2009). "Catalogue of the continental mollusks of Russia and adjacent territories". Version 2.3.
4. Páll-Gergely, Barna (2009). "REVISION OF THE TURKISH RAMUSCULUS TAXA WITH DESCRIPTION OF AYNA GEN. NOV. (GASTROPODA: PULMONATA: ENIDAE)" 40 (1): 73–77.
5. Schileyko A. A. (2007). "On the genus Mastoides Westerlund, 1896 (Pulmonata, Enidae)". Ruthenica 17(1-2): 85-87. abstract.
6. Hausdorf H. (1999) "A new genus of the Buliminidae from Turkey (Gastropoda: S tylommatophora)" Journal of Natural History 33: 149-154. PDF
7. Altaba C. R. (2007). "A new genus and species of Enidae (Gastopoda (sic!): Pulmonata) from the Quaternary of the Balearic Islands (Western Mediterranean)". Zootaxa 1595: 43-52. first page only.

External links

• "Enidae". National Center for Biotechnology Information (NCBI).