Bela (gastropod)

Bela
Shell of Bela nebula, type species of the genus Bela
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
(unranked):	clade Caenogastropoda clade Hypsogastropoda clade Neogastropoda
Superfamily:	Conoidea
Family:	Mangeliidae [1]
Genus:	Bela J. E. Gray, 1847[2]
Type species
Bela nebula (Montagu, 1803)
Synonyms[3]
Fehria van Aartsen, 1988 Ginnania Monterosato, 1884 Mangelia (Bela) J. E. Gray, 1847

Bela is a genus of sea snails; marine gastropod mollusks in the family Mangeliidae.^[3]

Species

Species within the genus Bela (gastropod) include:

• Bela africana Ardovini, 2004^[4]
• Bela alma Thiele, 1925^[5]
• Bela anderssoni Strebel, 1908^[6]
• Bela anna Thiele, 1925^[7]
• Bela ardovinii Mariottini & Oliverio, 2008^[8]
• Bela bella Barnard, 1958^[9]
• Bela brachystoma (Philippi, 1844)^[10]
• Bela chuni Thiele, 1925^[11]
• Bela clarae Peñas & Rolán, 2008^[12]
• Bela cycladensis (Reeve, 1845)^[13]
• Bela decussata (Locard, 1897)^[14]
• Bela erosa (Schrenck, 1861)
• Bela exilis Ardovini, 2004^[15]
• Bela fulvicans Strebel, 1908^[16]
• Bela fuscata (Deshayes, 1835)^[17]
• Bela glacialis Thiele, 1912^[18]
• Bela menkhorsti van Aartsen, 1988^[19]
• Bela nebula (Montagu, 1803)^[20]
• Bela oceanica (Locard, 1897)^[21]
• Bela powisiana (Dautzenberg, 1887)^[22]
• Bela taprurensis (Pallary, 1904)
• † Bela trinacria Mariottini & Smriglio, 2009^[23]
• Bela zenetouae (van Aartsen, 1988)^[24]
• Bela zonata (Locard, 1892)^[25]

Species brought into synonymy 

• Bela abyssorum Locard, 1897 : synonym of Gymnobela abyssorum (Locard, 1897)
• Bela aegeensis (Reeve, 1844):^[26] synonym of Bela nebula (Montagu, 1803)
• Bela alberti (Dautzenberg & Fisher, 1906):^[27] synonym of Phymorhynchus alberti (Dautzenberg & Fischer, 1906)
• Bela albrechti Krause, 1886 : synonym of Granotoma albrechti (Krause, 1886)
• Bela angulosa Sars G. O., 1878 : synonym of Propebela angulosa (G. O. Sars, 1878)
• Bela antarctica Strebel, 1908 : synonym of Conorbela antarctica (Strebel, 1908)
• Bela assimilis Sars G. O., 1878 : synonym of Propebela assimilis (Sars G. O., 1878)
• Bela australis Adams & Angas, 1864 : synonym of Guraleus australis (Adams & Angas, 1864)
• Bela bergensis Friele, 1886 : synonym of Propebela bergensis (Friele, 1886)
• Bela blaneyi Bush, 1909:^[28] synonym of Oenopota blaneyi (Bush, 1909)
• Bela concinnula Verrill, 1882 : synonym of Propebela concinnula (A. E. Verrill, 1882)
• Bela confusa (Locard, 1897):^[29] synonym of Bela brachystoma (Philippi, 1844)
• Bela conoidea Sars G. O., 1878 : synonym of Curtitoma conoidea (Sars G. O., 1878)
• Bela costulata (Risso, 1826):^[30] synonym of Mangelia costulata Risso, 1826
• Bela eva Thiele, 1925 : synonym of Maoritomella eva (Thiele, 1925)
• Bela excurvata Carpenter, 1864 : synonym of Oenopota excurvata (Carpenter, 1864)
• Bela expansa Sars G. O., 1878 : synonym of Lusitanops expansa (Sars G. O., 1878)
• Bela graphica Locard, 1897 : synonym of Oenopota graphica (Locard, 1897)
• Bela ginnania (Risso, 1826):^[26] synonym of Bela nebula
• Bela grimaldii Dautzenberg, 1889 : synonym of Amphissa acutecostata (Philippi, 1844)
• Bela grippi Dall, 1908 : synonym of Bellaspira grippi (Dall, 1908)
• Bela guernei Dautzenberg, 1891 : synonym of Belomitra quadruplex (Watson, 1882)
• Bela harpa Dall, 1885 : synonym of Oenopota harpa (Dall, 1885)
• Bela hebes Verrill, 1880 : synonym of Curtitoma hebes (Verrill, 1880)
• Bela iessoensis Smith E. A., 1875 : synonym of Obesotoma iessoensis (Smith E. A., 1875)
• Bela incisula Verrill, 1882 : synonym of Curtitoma incisula (Verrill, 1882)
• Bela koreni Friele, 1886 : synonym of Oenopota koreni (Friele, 1886)
• Bela krausei Dall, 1887 : synonym of Granotoma krausei (Dall, 1887)
• Bela laevigata (Philippi, 1836):^[31] synonym of Bela zonata (Locard, 1892)
• Bela limatula Locard, 1896 : synonym of Amphissa acutecostata (Philippi, 1844)
• Bela martensi Strebel, 1905 : synonym of Mangelia martensi (Strebel, 1905)
• Bela metschigmensis Krause, 1886 : synonym of Oenopota metschigmensis (Krause, 1886)
• Bela mingoranceae Martin Perez & Vera-Pelaez, 2006:^[32] synonym of Bela powisiana (Dautzenberg, 1887)
• Bela mitralis Adams & Angas, 1864 : synonym of Guraleus mitralis (Adams & Angas, 1864)
• Bela murdochiana Dall, 1885 : synonym of Oenopota murdochiana (Dall, 1885)
• Bela neozelanica Suter, 1908 : synonym of Scrinium neozelanica (Suter, 1908)
• Bela notophila Strebel, 1908 : synonym of Lorabela notophila (Strebel, 1908)
• Bela obliqua Sars G. O., 1878 : synonym of Oenopota obliqua (Sars G.O., 1878)
• Bela optima Thiele, 1925 : synonym of Microdrillia optima (Thiele, 1925)
• Bela ornata (Locard, 1891):^[33] synonym of Mangelia costulata Risso, 1826
• Bela pelseneri Strebel, 1908 : synonym of Lorabela pelseneri (Strebel, 1908)
• Bela plicatula Thiele, 1912 : synonym of Lorabela plicatula (Thiele, 1912)
• Bela purissima Strebel, 1908 : synonym of Typhlodaphne purissima (Strebel, 1908)
• Bela rathbuni Verrill, 1882 : synonym of Propebela rathbuni (Verrill, 1882)
• Bela regina Thiele, 1925 : synonym of Tomopleura regina (Thiele, 1925)
• Bela robusta Packard, 1866 : synonym of Obesotoma robusta (Packard, 1866)
• Bela sansibarica Thiele, 1925 : synonym of Microdrillia sansibarica (Thiele, 1925)
• Bela scalaroides Sars G. O., 1878 : synonym of Propebela scalaroides (Sars G. O., 1878)
• Bela sculpturata Dall, 1887 : synonym of Mangelia sculpturata (Dall, 1887)
• Bela septenvillei (Dautzenberg & Durouchoux, 1913):^[34] synonym of Bela nebula (Montagu, 1803)
• Bela solida Dall, 1887 : synonym of Obesotoma solida (Dall, 1887)
• Bela spitzbergensis Friele, 1886 : synonym of Propebela spitzbergensis (Friele, 1886)
• Bela striatula Thiele, 1912 : synonym of Belalora striatula (Thiele, 1912)
• Bela subturgida Verrill, 1884 : synonym of Propebela subturgida (Verrill, 1884)
• Bela subvitrea Verrill, 1884 : synonym of Propebela subvitrea (Verrill, 1884)
• Bela tenuicostata Sars G. O., 1878 : synonym of Oenopota tenuicostata (Sars G.O., 1878)
• Bela tenuilirata Dall, 1871 : synonym of Obesotoma tenuilirata (Dall, 1871)
• Bela tumida Posselt, 1898 : synonym of Obesotoma tumida (Posselt, 1898)
• Bela turrita Strebel, 1908 : synonym of Belaturricula turrita (Strebel, 1908)
• Bela yanamii Yokoyama, 1926 : synonym of Plicifusus yanamii (Yokoyama, 1926)
• Bela zonatum (Locard, 1891):^[35] duplicate of Bela zonata

Nomina dubia 

• Bela fortis (Reeve, 1844):^[36] nomen dubium
• Bela minuta (Reeve, 1844):^[37] nomen dubium
• Bela turgida (Reeve, 1844):^[38] nomen dubium

References

1. Bouchet P., Kantor Yu.I., Sysoev A. & Puillandre N. (2011) A new operational classification of the Conoidea. Journal of Molluscan Studies 77: 273-308.
2. Gray J. E. (1847). Ann. Mag. nat. Hist. 20: 270.
3. Bela (gastropod) Gray, 1847.  Retrieved through: World Register of Marine Species on 18 February 2011.
4. Bela africana Ardovini, 2004.  Retrieved through: World Register of Marine Species on 16 April 2010.
5. Bela alma Thiele, 1925.  Retrieved through: World Register of Marine Species on 16 April 2010.
6. Bela anderssoni Strebel, 1908.  Retrieved through: World Register of Marine Species on 16 April 2010.
7. Bela anna Thiele, 1925.  Retrieved through: World Register of Marine Species on 16 April 2010.
8. Bela ardovinii Mariottini & Oliverio, 2008.  Retrieved through: World Register of Marine Species on 16 April 2010.
9. Bela bella Barnard, 1958.  Retrieved through: World Register of Marine Species on 16 April 2010.
10. Bela brachystoma (Philippi, 1844).  Retrieved through: World Register of Marine Species on 16 April 2010.
11. Bela chuni Thiele, 1925.  Retrieved through: World Register of Marine Species on 16 April 2010.
12. Bela clarae Peñas & Rolán, 2008.  Retrieved through: World Register of Marine Species on 16 April 2010.
13. Bela cycladensis (Reeve, 1845).  Retrieved through: World Register of Marine Species on 16 April 2010.
14. Bela decussata (Locard, 1897).  Retrieved through: World Register of Marine Species on 16 April 2010.
15. Bela exilis Ardovini, 2004.  Retrieved through: World Register of Marine Species on 16 April 2010.
16. Bela fulvicans Strebel, 1908.  Retrieved through: World Register of Marine Species on 16 April 2010.
17. Bela fuscata (Deshayes, 1835).  Retrieved through: World Register of Marine Species on 16 April 2010.
18. Bela glacialis Thiele, 1912.  Retrieved through: World Register of Marine Species on 16 April 2010.
19. Bela menkhorsti van Aartsen, 1988.  Retrieved through: World Register of Marine Species on 16 April 2010.
20. Bela nebula (Montagu, 1803).  Retrieved through: World Register of Marine Species on 16 April 2010.
21. Bela oceanica (Locard, 1897).  Retrieved through: World Register of Marine Species on 16 April 2010.
22. Bela powisiana (Dautzenberg, 1887).  Retrieved through: World Register of Marine Species on 16 April 2010.
23. Mariottini P., Smriglio C., Di Giulio A. & Oliverio M. (2009). "A new fossil conoidean from the Pliocene of Italy, with comments on the Bela menkhorsti complex (Gastropoda: Conidae)". Journal of Conchology 40(1): 5-14. abstract.
24. Bela zenetouae (van Aartsen, 1988).  Retrieved through: World Register of Marine Species on 16 April 2010.
25. Bela zonata (Locard, 1892).  Retrieved through: World Register of Marine Species on 16 April 2010.
26. B. Otztürk, Comments on Shells of Mollusca collected from the Seas of Turkey, Turk J Zool, 29, pp. 111-112, 2005; accessed : 20 February 2011
27. Bela alberti (Dautzenberg & Fisher, 1906).  Retrieved through: World Register of Marine Species on 16 April 2010.
28. Bela blaneyi Bush, 1909.  Retrieved through: World Register of Marine Species on 16 April 2010.
29. Bela confusum (Locard, 1897).  Retrieved through: World Register of Marine Species on 16 April 2010.
30. Bela costulata (Risso, 1826).  Retrieved through: World Register of Marine Species on 16 April 2010.
31. Bela laevigata (Philippi, 1836).  Retrieved through: World Register of Marine Species on 16 April 2010.
32. Bela mingoranceae Martin Perez & Vera-Pelaez, 2006.  Retrieved through: World Register of Marine Species on 16 April 2010.
33. Bela ornata (Locard, 1891).  Retrieved through: World Register of Marine Species on 16 April 2010.
34. Bela septenvillei (Dautzenberg & Durouchoux, 1913).  Retrieved through: World Register of Marine Species on 16 April 2010.
35. Bela zonatum (Locard, 1891).  Retrieved through: World Register of Marine Species on 16 April 2010.
36. Bela fortis (Reeve, 1844).  Retrieved through: World Register of Marine Species on 16 April 2010.
37. Bela minuta (Reeve, 1844).  Retrieved through: World Register of Marine Species on 16 April 2010.
38. Bela turgida (Reeve, 1844).  Retrieved through: World Register of Marine Species on 16 April 2010.

• Vaught, K.C. (1989). A classification of the living Mollusca. American Malacologists: Melbourne, FL (USA). ISBN 0-915826-22-4. XII, 195 pp
• Howson, C.M.; Picton, B.E. (Ed.) (1997). The species directory of the marine fauna and flora of the British Isles and surrounding seas. Ulster Museum Publication, 276. The Ulster Museum: Belfast, UK. ISBN 0-948150-06-8. vi, 508
• Gofas, S.; Le Renard, J.; Bouchet, P. (2001). Mollusca, in: Costello, M.J. et al. (Ed.) (2001). European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels, 50: pp. 180–213
• Mariottini P., Smriglio C., Di Giulio A. & Oliverio M. 2009. A new fossil conoidean from the Pliocene of Italy, with comments on the Bela menkhorsti complex (Gastropoda: Conidae). Journal of Conchology 40(1): 5-14

External links