Glyoxysome

Glyoxysomes are specialized peroxisomes found in plants (particularly in the fat storage tissues of germinating seeds) and also in filamentous fungi. As in all peroxisomes, in glyoxysomes the fatty acids are hydrolyzed to acetyl-CoA by peroxisomal β-oxidation enzymes. Besides peroxisomal functions, glyoxysomes possess additionally the key enzymes of glyoxylate cycle (isocitrate lyase and malate synthase) which accomplish the glyoxylate cycle bypass.

Thus, glyoxysomes (as all peroxisomes) contain enzymes that initiate the breakdown of fatty acids and additionally possess the enzymes to produce intermediate products for the synthesis of sugars by gluconeogenesis. The seedling uses these sugars synthesized from fats until it is mature enough to produce them by photosynthesis.

Glyoxysomes also participate in photorespiration and nitrogen metabolism in root nodules.

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MeSH A11.284.195.190.500.585.250

Structures of the cell / organelles (TH H1.00.01.2-3)

Endomembrane system	Cell membrane Nucleus Nucleolus Endoplasmic reticulum Golgi apparatus Parenthesome Autophagosome Vesicles Exosome Lysosome Endosome Phagosome Vacuole Acrosome Cytoplasmic granules Melanosome Microbody Glyoxysome Peroxisome Weibel–Palade body

Cytoskeleton	Microfilaments Intermediate filaments Microtubules Prokaryotic cytoskeleton MTOCs Centrosome Centriole Basal body Spindle pole body Myofibril

Endosymbionts	Mitochondrion Plastids Chloroplast Chromoplast Gerontoplast Leucoplast Amyloplast Elaioplast Proteinoplast Tannosome

Other internal	RNA Ribosome Spliceosome Vault Cytoplasm Proteasome

External	Undulipodium Cilium Flagellum Axoneme Radial spoke Extracellular matrix Cell wall

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