Hadrianus (genus)

Hadrianus Temporal range: 55.4–33.9 Ma PreЄ Є O S D C P T J K Pg N Eocene^[1]
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Family:	Testudinidae
Genus:	†Hadrianus Cope, 1872^[1]
Species
Hadrianus corsoni (syn. H. octonarius) Hadrianus robustus Hadrianus schucherti Hadrianus utahensis

Hadrianus is an extinct genus of tortoise belonging to the Testudinidae^[1] found in the United States and Spain and believed to be the oldest true tortoise known.^[2] The genus is thought to be closely related to the genus Manouria.^[2] The genus may have evolved in the subtropics of Asia and subsequently migrated to North America and Europe.^[2]

References

1 2 3 http://paleodb.org/cgi-bin/bridge.pl?action=checkTaxonInfo&taxon_no=58498&is_real_user=1
1 2 3 Ehret, Dana Joseph 2004 "Skeletochronology as a method of aging Oligocene Gopherus laticuneus and Stylemys nebrascensis, using Gopherus polyphemus as a modern analog" Thesis, University of Florida.

Extinct turtles

Triassic period	Pappochelys Odontochelys Proganochelys

Jurassic period	Australochelys Basilochelys Caribemys Condorchelys Dinochelys Dorsetochelys Eileanchelys Eurysternum Hispaniachelys Kayentachelys Palaeomedusa Sinaspideretes Thalassemys Uluops

Cretaceous period	Allopleuron Atlantochelys Angolachelys Anomalochelys Araripemys Archelon Bouliachelys Caririemys Cearachelys Chelosphargis Chupacabrachelys Compsemys Corsochelys Cratochelone Ctenochelys Desmatochelys Emarginachelys Foxemys Gigantatypus Gilmoremys Glyptops Jiangxichelys Kallokibotion Kinkonychelys Kirgizemys Khunnuchelys Kurmademys Liaochelys Lomalatachelys Notochelone Ocepechelon Pappigerus Paracryptodira Patagoniaemys Polysternon Pleurosternon Prochelidella Protostega Protostegidae Pneumatoarthrus Sankuchemys Santanachelys Sinemys Terlinguachelys Toxochelys Trionyx kansaiensis Yaminuechelys

Paleogene and Neogene period	Acherontemys Allaeochelys Anosteira Arabemys Argillochelys Aurorachelys Baenidae Berruchelus Carbonemys Carolinochelys Cedrobaena Cerrejonemys Chelodina murrayi Chelydrops Chelydropsis Chisternon Cosmochelys Crossochelys Elseya uberrima Elseya nadibajagu Eochelone Eosphargis Euclastes Gamerabaena Glarichelys Hadrianus Hesperotestudo Hutchemys Itilochelys Kinosternon pojoaque Macrocephalochelys Manchurochelys Meiolania Meiolaniidae Megalochelys Mesodermochelys Mexichelys Miocaretta Niolamia Ninjemys Pacifichelys Palaeotrionyx Palatobaena Parahydraspis Peckemys Planiplastron Plesiobaena Procolpochelys Protochelydra Protosphargis Psephophorus Puppigerus Rheodytes devisi Stupendemys Stylemys Syllomus Warkalania

Recent	Cylindraspis species: Domed Mauritius giant tortoise Domed Rodrigues giant tortoise Réunion giant tortoise Saddle-backed Mauritius giant tortoise Saddle-backed Rodrigues giant tortoise Aldabrachelys abrupta Chelonoidis nigra abingdonii (Lonesome George)

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