Hadrosauroidea

Hadrosauroids Temporal range: Early-Late Cretaceous, 130–66 Ma PreЄ Є O S D C P T J K Pg N
Holotype skeleton of Tethyshadros insularis
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Order:	†Ornithischia
Suborder:	†Ornithopoda
Clade:	†Hadrosauriformes
Superfamily:	†Hadrosauroidea Cope, 1869
Type species
†Hadrosaurus foulkii Leidy, 1858
Families
†Hadrosauridae And see text.
Synonyms
Trachodontoidea Lydekker, 1888 Hadrosauria Huene, 1956

Hadrosauroidea is a clade or superfamily of ornithischian dinosaurs that includes the "duck-billed" dinosaurs, or hadrosaurids, and all dinosaurs more closely related to them than to Iguanodon. Many primitive hadrosauroids, such as the sail-backed Ouranosaurus, have traditionally been included in a paraphyletic (unnatural grouping) "Iguanodontidae". With cladistic analysis, the traditional Iguanodontidae has been largely disbanded, and probably includes only Iguanodon and perhaps its closest relatives.

Classification

Taxonomy

The following taxonomy follows Ramírez-Velasco et al, 2012 unless otherwise noted.^[1]

• Superfamily Hadrosauroidea
  • Altirhinus^[2]
  • Bactrosaurus
  • Batyrosaurus^[3]
  • Bolong^[3]
  • Claosaurus
  • Equijubus
  • Eolambia
  • Fukuisaurus
  • Gilmoreosaurus
  • Glishades
  • Huehuecanauhtlus
  • Jeyawati
  • Jintasaurus
  • Jinzhousaurus
  • Koshisaurus^[4]
  • Levnesovia
  • Lophorhothon
  • Nanyangosaurus
  • Penelopognathus
  • Probactrosaurus
  • Protohadros
  • Siamodon
  • Shuangmiaosaurus
  • Tanius
  • Tethyshadros
  • Xuwulong^[5]
  • Family Hadrosauridae

Phylogeny

The cladogram below follows an analysis by Wu Wenhao and Pascal Godefroit (2012).^[3]

Hadrosauriformes	Iguanodontidae  Iguanodon Ouranosaurus Mantellisaurus  Hadrosauroidea  Bolong Equijubus Jinzhousaurus Altirhinus Batyrosaurus Probactrosaurus Eolambia Protohadros Bactrosaurus Levnesovia Shuangmiaosaurus Tethyshadros Telmatosaurus Hadrosauridae

Cladogram after Prieto-Marquez and Norell (2010).^[6]

Hadrosauroidea	Jinzhousaurus Penelopognathus Equijubus Probactrosaurus Eolambia Protohadros Tanius Bactrosaurus Gilmoreosaurus Telmatosaurus Shuangmiaosaurus Nanyangosaurus Lophorhothon Hadrosauridae

A phylogenetic analysis performed by Ramírez-Velasco et al. (2012) found a big polytomy of all hadrosauroids which are more derived than Probactrosaurus but less derived than Hadrosauridae. The exclusion of Claosaurus, Jeyawati, Levnesovia, Nanyangosaurus, Shuangmiaosaurus and Telmatosaurus from the polytomy resulted in a more resolved topology. The cladogram below shows Huehuecanauhtlus phylogenetic position among other hadrosauroids following this topology (the relationships within Hadrosauridae are not shown).^[1]

Hadrosauroidea	Jinzhousaurus yangi Fukuisaurus tetoriensis Penelopognathus weishampeli Equijubus normani Probactrosaurus gobiensis Eolambia caroljonesa Protohadros byrdi Tanius sinensis Bactrosaurus johnsoni Glishades ericksoni Gilmoreosaurus mongoliensis Huehuecanauhtlus tiquichensis Jintasaurus meniscus Tethyshadros insularis Hadrosauridae

See also

• Timeline of hadrosaur research

References

1. Angel Alejandro Ramírez-Velasco, Mouloud Benammi, Albert Prieto-Márquez, Jesús Alvarado Ortega and René Hernández-Rivera (2012). "Huehuecanauhtlus tiquichensis, a new hadrosauroid dinosaur (Ornithischia: Ornithopoda) from the Santonian (Late Cretaceous) of Michoacán, Mexico". Canadian Journal of Earth Sciences 49 (2): 379–395. doi:10.1139/e11-062.
2. Hai-Lu You; Da-Qing Li (2009). "A new basal hadrosauriform dinosaur (Ornithischia: Iguanodontia) from the Early Cretaceous of northwestern China". Canadian Journal of Earth Sciences 46 (12): 949–957. doi:10.1139/E09-067.
3. Wu Wenhao and Pascal Godefroit (2012). "Anatomy and Relationships of Bolong yixianensis, an Early Cretaceous Iguanodontoid Dinosaur from Western Liaoning, China". In Godefroit, P. (eds). Bernissart Dinosaurs and Early Cretaceous Terrestrial Ecosystems. Indiana University Press. pp. 293–333.
4. Shibaba, Masateru; Azuma, Yoichi (2015). "New basal hadrosauroid (Dinosauria: Ornithopoda) from the Lower Cretaceous Kitadani Formation, Fukui, central Japan." (PDF). Zootaxa 3914: 421. doi:10.11646/zootaxa.3914.4.3.|
5. McDonald, A. T. (2012). Farke, Andrew A, ed. "Phylogeny of Basal Iguanodonts (Dinosauria: Ornithischia): An Update". PLoS ONE 7 (5): e36745. doi:10.1371/journal.pone.0036745. PMC 3358318. PMID 22629328.
6. Albert Prieto-Marquez and Mark A. Norell (2010). "Anatomy and Relationships of Gilmoreosaurus mongoliensis (Dinosauria: Hadrosauroidea) from the Late Cretaceous of Central Asia" (PDF). American Museum Novitates 3694: 1–52. doi:10.1206/3694.2. ISSN 0003-0082.