Andamanese people

Two Great Andamanese men, circa. 1875

The Andamanese people are the various aboriginal inhabitants of the Andaman Islands, a Union Territory of India located in the southeastern part of the Bay of Bengal.

The Andamanese resemble other Negrito groups in Southeast Asia. They are pygmies, and are the only modern people outside of certain parts of Sub-Saharan Africa with steatopygia. They lead a hunter-gatherer lifestyle, and appear to have lived in substantial isolation for thousands of years. The Andamanese arrived at the Andaman Islands around the latest Glacial Maximum, ca. 26,000 years ago.

By the end of the eighteenth century, when they first came into sustained contact with outsiders, there were an estimated 7,000 Andamanese divided into five major groups, with distinct cultures, separate domains, and mutually unintelligible languages. In the next century they were largely wiped out by diseases, violence, and loss of territory. Today, there remain only approximately 400–450 Andamanese. One group has long been extinct, and only two of the remaining groups still maintain a steadfast independence, refusing most attempts at contact by outsiders.

The Andamanese are a designated Scheduled Tribe.[1]

History

Until the late eighteenth century, the Andamanese culture, language, and genetics were preserved from outside influences by their fierce reaction to visitors, which included killing any shipwrecked foreigners, and by the remoteness of the islands. The various tribes and their mutually unintelligible languages thus are believed to have evolved on their own over millennia.

Origins

Map of human haplotype migration based on mitochondrial DNA, with Key (coloured) indicating periods in numbered thousands of years before the present. Note the route of the mtDNA haplogroup M through the Indian mainland and the Andaman Islands, possibly on to Southeast Asia

According to Chaubey and Endicott (2013), the Andaman Islands were settled less than 26,000 years ago, by people who were not direct descendants of the first migrants out of Africa.[2][note 1] According to Wang et al. (2011),

...the Andaman archipelago was likely settled by modern humans from northeast India via the land-bridge which connected the Andaman archipelago and Myanmar around the Last Glacial Maximum (LGM), a scenario in well agreement with the evidence from linguistic and palaeoclimate studies.[3]

It was previously assumed that the Andaman ancestors were part of the initial Great Coastal Migration that was the first expansion of humanity out of Africa, via the Arabian peninsula, along the coastal regions of the Indian mainland and toward Southeast Asia, Japan, and Oceania.[4][5] The Andamese were considered to be a pristine example of a hypothised Negrito population, which showed similar physical characteristics, and was supposed to have existed throughout southeast Asia. The existence of a specific Negrito-population is nowadays doubted. Their commonalities could be the result of evolutionary convergence and/or a shared history.[6][7]

Population decline

An official 1867 British government communication about organizing a punitive expedition against Andamanese tribespeople on Little Andaman Island

The Andamanese's protective isolation changed with the first British colonial presence and subsequent settlements, which proved disastrous for them. Lacking immunity against common diseases of the Eurasian mainland, the large Jarawa habitats on the southeastern regions of South Andaman Island likely were depopulated by disease within four years (1789-1793) of the initial British colonial settlement in 1789.[8] Epidemics of pneumonia, measles and influenza spread rapidly and exacted heavy tolls, as did alcoholism.[8] By 1875, the Andamanese were already "perilously close to extinction," yet attempts to contact, subdue and co-opt them continued unrelentingly. In 1888, the British government set in place a policy of "organized gift giving" that continued in varying forms until well into the 20th century.[9]

Great Andamese women, 1876

There is evidence that some sections of the British Indian administration were working deliberately to annihilate the tribes.[10] After the mid-19th century, British established penal colonies on the islands and an increasing numbers of mainland Indian and Karen settlers arrived, encroaching on former territories of the Andamanese. This accelerated the decline of the tribes.

Many Andamanese succumbed to British expeditions to avenge the killing of shipwrecked sailors. In the 1867 Andaman Islands Expedition, dozens of Onge were killed by British naval personnel following the death of shipwrecked sailors, which resulted in four Victoria Crosses for the British soldiers.[11][12][13] In the 1940s, the Jarawa were bombed by Japanese forces for their hostility.[14]

Recent history

In 1974, a film crew and anthropologist Triloknath Pandit attempted friendly contact by leaving a tethered pig, some pots and pans, some fruit and toys on the beach at North Sentinel Island. One of the islanders shot the film director in the thigh with an arrow. The following year, European visitors were repulsed with arrows.[15][16][17]

On 2 August 1981, the Hong Kong freighter ship Primrose grounded on the North Sentinel Island reef. A few days later, crewmen on the immobile vessel observed that small black men were carrying spears and arrows and building boats on the beach. The captain of the Primrose radioed for an urgent airdrop of firearms so the crew could defend themselves but did not receive them. Heavy seas kept the islanders away from the ship. After a week, the crew were rescued by an Indian navy helicopter.[18]

On 4 January 1991, Indian scholar Triloknath Pandit made the first known friendly contact with the Sentinelese.[17]

Until 1996, the Jarawa met most visitors with flying arrows. From time to time they attacked and killed poachers on the lands reserved to them by the Indian government. They also killed some workers building the Andaman Trunk Road (ATR), which traverses Jarawa lands. One of the earliest peaceful contacts with the Jarawa occurred in 1996. Settlers found a teenage Jarawa boy named Emmei near Kadamtala town. The boy was immobilized with a broken foot. They took Emmei to a hospital where he received good care. Over several weeks, Emmei learned a few words of Hindi before returning to his jungle home. The following year, Jarawa individuals and small groups began appearing along roadsides and occasionally venturing into settlements to steal food. The ATR may have interfered with traditional Jarawa food sources.[19][20][21]

Tribes

Distribution of Andamanese tribes in the Andaman Islands — early 1800s versus present-day (2004)

The five major groups of Andamanese found by the European colonists were:

Jarawa and Onge are dominated by y-dna haplogroup D which is native to the Andamanese.[23]

By the end of the eighteenth century, there were an estimated 5,000 Great Andamanese living on Great Andaman. Altogether they comprised ten distinct tribes with different languages. The population quickly dwindled until they became extinct in pure form, and had a low of 19 admixed individuals by 1961.[24] It has increased slowly after that, following their move to a reservation on Strait Island. By January 2011, there were only 54 admixed individuals from three tribes, who spoke mostly Hindi.[22]

The Jarawa originally inhabited southeastern Jarawa Island and have migrated to the west coast of Great Andaman in the wake of the Great Andamanese. The Onge once lived throughout Little Andaman and now are confined to two reservations on the island. The Jangil, who originally inhabited Rutland Island, were extinct by 1931; the last individual was sighted in 1907.[14] Only the Sentinelese are still living in their original homeland on North Sentinel Island, largely undisturbed, and have fiercely resisted all attempts at contact.

The tribe members of the Jarawa tribe focused on traditional knowledge. Traditional knowledge was a huge part of their culture. This referred to those who learned through observations and experiences when it came to identifying the relationship between different organisms and environmental resource areas. Their way of life is interwoven with the traditional knowledge of a group.[25]

Languages

Main article: Andamanese languages

The Andamanese languages are considered to be the fifth language family of India, following the Indo-European, Dravidian, Austroasiatic, and Sino-Tibetan.[26]

While some connections have been tentatively proposed with other language families,[27] the consensus view is currently that Andamanese languages form a separate language family — or rather, two unrelated linguistic families: Greater Andamanese on the one hand,[28] and Ongan on the other hand.

Culture

Group of Andamanese hunting turtles with bows and arrows

Until contact, the Andamanese were strict hunter-gatherers. They did not practice cultivation, and lived off hunting indigenous pigs, fishing, and gathering. Their only weapons were the bow, adzes, and wooden harpoons. Besides the aboriginal people of Tasmania, the Andamanese were the only people who in the nineteenth century knew no method of making fire. They instead carefully preserved embers in hollowed-out trees from fires caused by lightning strikes. They are known as Chadda.

The men wore girldes made of hibiscus fiber which carried useful tools and weapons for when they went hunting.The women on the other hand wore a tribal dress containing leaves that were held by a belt. Majority of them had painted bodies as well. They usually slept on leaves as well, or mats and had either permanent or temporary habitation among the tribes. All habitations were man built.[29] For hunting, different types of methods were used. They survived over thousands of years by hunting and gathering. They made spears, bows, arrows...etc., specific to what they were hunting. The designing of the various weapons and tools show how their levels of excellence in skill, craftsmanship and creativity were.[25]

Some of the tribe members were credited to having super natural powers. They were called oko-pai-ad, which meant dreamer. They were thought to have an influence on the members of the tribe and would bring misfortune to those who did not believed in their abilities. Traditional knowledge practitioners were the ones who helped with healthcare. The medicine that was used to cure illnesses were herbal most of the time. Various types of medicinal plants were used by the islanders.77 total traditional knowledge practitioners were identified and 132 medicinal plants were used. the members of the tribes found various ways to use leaves in their everyday lives including clothing, medicine, and to sleep on.[30]

Physical appearance

Phenotype

Negritos, specifically Andamanese, are grouped together by phenotype features. Three physical features that distinguish the Andaman islanders include: skin color, hair, and stature. Those of the Andaman islands have dark skin, are short in stature, and have "frizzy" hair. These physical features are shared with Melanesians and believed to have been retained from the early dispersing Homo sapiens. [31][32]

The frizzy hair is considered polygenic because it is the one phenotype feature that has not changed. The skin color and stature can vary within a population.[32] Therefore, hair characterization is very important in identifying the Andaman islanders. The stature of the people within the population has changed over time, a relatively short time. Scientists believe the variation in stature can be contributed to about 200 gene loci that influence height. Similarly, variation in skin color can be contributed to the great number of gene loci that affect the expression of that phenotype. The isolation of the Andamanese people is what has kept the variation in phenotype within the population low. The craniometric study of the Andamanese people is almost the same as that of the South Asians. Therefore, the only way to distinguish between South Asians and Andamanese people is by comparing their stature, skin color, and hair.

Dental Morphology

At first, the dental morphology of the Andamanese was in the middle between African and South Asian. Further study suggests that the dental morphology is Sundadont in pattern; therefore most similar to the dental morphology of South Asians.[33]

When comparing dental morphology the focus is on overall size and tooth shape. To measure the size and shape, Penrose's size and shape statistic is used. To calculate tooth size, the sum of the tooth area is taken. Factor analysis is applied to tooth size to achieve tooth shape. Results have shown that tooth size of Andaman islanders is closest to the tooth size of Chinese and Japanese people . Therefore, the indodont dental morphology of the Andamanese indicates a retention of dental morphology from Southeast Asians in early-mid Holocene.[31]

Skeletal Phenotype

The Andaman islanders are characterized by their small bodies and glacial skeleton.[34] Additionally they have a narrow bi-iliac breadth and short upper limbs. The craniometric studies suggest that the Andamanese are most similar to African in terms of the craniology. The small body size of the Andamanese results in a high surface area to mass ratio, low rate of metabolism, and less heat produced. This is a great adaptation in a hot environment.[33]

Genetics

"Scarification pattern among the Great Andamanese in the late 19th century. Nothing is known of the origins or antiquity of this custom among the Andamanese."

Genetic analysis, both of nuclear DNA[4][35] and mitochondrial DNA[36] provide information about the origins of the Andamese. The Andamanese are similar to Africans in terms of Craniology and are genetically closest to East Asians. The Andaman islanders have similar dental morphology to South Asians.[31] The Andamanese are most genetically similar to the Malaysian Negrito tribe.[37]

Nuclear DNA

The Andamese show a very small genetic variation, which is indicative of populations that have experienced a population bottleneck and then developed in isolation for a long period. An allele has been discovered among the Jarawas that is found nowhere else in the world. Overall, the Andamanese showed closest relations with other Asian populations. The Nicobarese also were observed to share close genetic relations with adjacent Indo-Mongoloid populations of Northeast India.[35] Bulbeck (2013) likewise noted that the Andamanese's nuclear DNA clusters with that of other Andamanese Islanders, as they carry Haplogroup D and maternal M (mtDNA) unique to their own.[31]

Genomic

The use of single-nucleotide polymorphism (SNP) shows the genomes of Andamanese people to be closest to those of South Asians. This suggests a relation between Andaman islanders and South Asians.[31] Analysis of mtDNA, which is inherited exclusively by maternal descent, confirms the above results. All Onge belong to M32 mtDNA, subgroup of M which is unique to Onge people.[36] While parental Y-DNA is exclusively Haplogroup D.[38]

All Andamanese belong to MtDNA M and Y-DNA Haplogroup D, which expanded at the late Pleistocene.[36][38][39]

Reich et al. (2009) reckon that the Andamanese split from mainland Asia 1,700 generations ago.[38] Andamanese are the only South Asian population in the study that lacked any Ancestral North Indian admixture.[38]

Y DNA

The male Y-chromosome in humans is inherited exclusively through paternal descent. Male Onges and Jarawas almost exclusively belong to Haplogroup D-M174.[40] The clade is most common today in Tibet and Japan, with its highest frequencies worldwide in the Pumi population of southwestern China (70.2%).[41] Haplogroup D-M174 also occurs frequently among the Ainu of Japan.[42] On the Indian mainland, it has been observed at low frequencies among Tibeto-Burman speakers.[40] Andamanese males were found to carry five different binary D haplotypes, all of which had previously been observed on the Indian subcontinent, in Southeast Asia and Melanesia.[43]

Male Great Andamanese do not have D-M174. The low resolution study discovered these clades as O, L, K, and P-M45, which means that Great Andamanese are extinct in pure form as all of their male lines have a post-1900 foreign origin and around half of their blood.[43]

Mitochondrial DNA

Analysis of mtDNA, which is inherited exclusively by maternal descent, confirms the above results. All Andamanese belong to M31 and M32 mtDNA, subgroup of M which is unique to Andamanese people.[36][31] The analysis of 20 coding regions in 20 samples of ancient Andamanese people and 12 samples of modern Indian populations changed the topology of the two lineages in South Asians. The data received suggests an M31a lineage in South Asians. This supports the genetic connection between South Asians and Andamanese people, which dates back to about 30kya.[44] Other mainland specific subgroup of M is distributed in the Asia, where it represents 60% of all maternal lineages.[43][45][46] According to Endicott et al (2002), this haplogroup originated with the earliest settlers of India during the coastal migration that brought the ancestors of the Andamanese to the Indian mainland, the Andaman Islands, and farther afield to Southeast Asia.[47]

Lack of Denisovan heritage

Unlike some Negrito populations of Southeast Asia, Andaman Islanders have been found to have no Denisovan ancestry.[48]

See also

Notes

  1. ↑ Chaubey and Endicott (2013):[2]
    * "these estimates suggest that the Andamans were settled less than ~26 ka and that differentiation between the ancestors of the Onge and Great Andamanese commenced in the Terminal Pleistocene." (p.167)
    * "In conclusion, we find no support for the settlement of the Andaman Islands by a population descending from the initial out-of-Africa migration of humans, or their immediate descendants in South Asia. It is clear that, overall, the Onge are more closely related to Southeast Asians than they are to present-day South Asians." (p.167)

References

  1. ↑ "List of notified Scheduled Tribes" (PDF). Census India. p. 27. Retrieved 15 December 2013.
  2. 1 2 Chaubey and Endicott (2013), The Andaman Islanders in a Regional Genetic Context: Reexamining the Evidence for an Early Peopling of the Archipelago from South Asia, Human Biology 85 (1-3)
  3. ↑ Wang, Hua-Wei; Mitra, Bikash; Chaudhuri, Tapas Kumar; Palanichamy, Malliya gounder; Kong, Qing-Peng; Zhang, Ya-Ping (2011-03-20). "Mitochondrial DNA evidence supports northeast Indian origin of the aboriginal Andamanese in the Late Paleolithic". Journal of Genetics and Genomics 38 (3): 117–122. doi:10.1016/j.jgg.2011.02.005.
  4. 1 2 Spencer Wells (2002), The Journey of Man: A Genetic Odyssey, Princeton University Press, ISBN 0-691-11532-X, ... the population of south-east Asia prior to 6000 years ago was composed largely of groups of hunter-gatherers very similar to modern Negritos ... So, both the Y-chromosome and the mtDNA paint a clear picture of a coastal leap from Africa to south-east Asia, and onward to Australia ... DNA has given us a glimpse of the voyage, which almost certainly followed a coastal route va India ...
  5. ↑ Anvita Abbi (2006), Endangered Languages of the Andaman Islands, Lincom Europa, ... to Myanmar by a land bridge during the ice ages, and it is possible that the ancestors of the Andamanese reached the islands without crossing the sea ... The latest figure in 2005 is 50 in all ...
  6. ↑ Jinam, Timothy A.; Phipps, Maude E.; Saitou, Naruya (2013-06-01). "Admixture Patterns and Genetic Differentiation in Negrito Groups from West Malaysia Estimated from Genome-wide SNP Data". Human Biology 85 (1-3): 173–188. doi:10.3378/027.085.0308. ISSN 0018-7143.
  7. ↑ Stock, Jay T. (2013-01-01). "The Skeletal Phenotype of "Negritos" from the Andaman Islands and Philippines Relative to Global Variation among Hunter-Gatherers". Human Biology 85 (1): 67–94. doi:10.3378/027.085.0304. ISSN 1534-6617. PMID 24297221.
  8. 1 2 Sita Venkateswar (2004), Development and Ethnocide: Colonial Practices in the Andaman Islands, IWGIA, ISBN 87-91563-04-6, As I have suggested previously, it is probable that some disease was introduced among the coastal groups by Lieutenant Colebrooke and Blair's first settlement in 1789, resulting in a marked reduction of their population. The four years that the British occupied their initial site on the south-east of South Andaman were sufficient to have decimated the coastal populations of the groups referred to as Jarawa by the Aka-bea-da.
  9. ↑ Richard B. Lee, Richard Heywood Daly (1999), The Cambridge Encyclopedia of Hunters and Gatherers, Cambridge University Press, ISBN 0-521-57109-X, By 1875, when these peoples were perilously close to extinction, the Andaman cultures came under scientific scrutiny ... In 1888, 'friendly relations' were established with Ongees through organized gift giving contacts ... As recently as 1985—92, government contacts have been initiated with Jarawas and Sentinelese through gift-giving, a contact procedure much like that carried out during British rule.
  10. ↑ Luigi Luca Cavalli-Sforza, Francesco Cavalli-Sforza (1995), The Great Human Diasporas: The History of Diversity and Evolution, Basic Books, ISBN 0-201-44231-0, Contact with whites, and the British in particular, has virtually destroyed them. Illness, alcohol, and the will of the colonials all played their part; the British governor of the time mentions in his diary that he received instructions to destroy them with alcohol and opium. He succeeded completely with one group. The others reacted violently.
  11. ↑ Madhusree Mukerjee (2003), The Land of Naked People, Houghton Mifflin Books, ISBN 0-618-19736-2, In 1927 Egon Freiherr von Eickstedt, a German anthropologist, found that around one hundred Great Andamanese survived, 'in dirty, half-closed huts, which primarily contain cheap European household effects'.
  12. ↑ The London Gazette: no. 23333. p. 6878. 17 December 1867. Retrieved 2008-08-23.
  13. ↑ Laxman Prasad Mathur (2003), Kala Pani: History of Andaman & Nicobar Islands, with a Study of Indiaʼs Freedom Struggle, Eastern Book Corporation, Snippet: "Immediately afterwards in another visit to Little Andaman to trace the sailors of a ship named 'Assam Valley' wrecked on its coast, Homfray's party was attacked by a large group of Onges."
  14. 1 2 3 George van Driem (2001), Languages of the Himalayas: An Ethnolinguistic Handbook of the Greater Himalayan Region: Containing an Introduction to the Symbiotic Theory of Language, BRILL, ISBN 90-04-12062-9, ... The Aka-Kol tribe of Middle Andaman became extinct by 1921. The Oko-Juwoi of Middle Andaman and the Aka-Bea of South Andaman and Rutland Island were extinct by 1931. The Akar-Bale of Ritchie's Archipelago, the Aka-Kede of Middle Andaman and the A-Pucikwar of South Andaman Island soon followed. By 1951, the census counted a total of only 23 Greater Andamanese and 10 Sentinelese. That means that just ten men, twelve women and one child remained of the Aka-Kora, Aka-Cari and Aka-Jeru tribes of Greater Andaman and only ten natives of North Sentinel Island ...
  15. ↑ Goodheart, Adam (Autumn 2000). "The Last Island of the Savages". The American Scholar 69 (4). JSTOR 41213066.
  16. ↑ Pandit
  17. 1 2 "Islanders running out of isolation: Tim McGirk in the Andaman Islands reports on the fate of the Sentinelese". The Independent (London). 10 January 1993.
  18. ↑ "Grounding". neatorama.com.
  19. ↑ Seksharia, Pankaj. "Jarawa excursions". frontline.in. Front Line. Retrieved 30 May 2015.
  20. ↑ Valley, Paul. "Under threat: an ancient tribe emerging from the forests". listserv.linguist.org. The Independent UK. Retrieved 30 May 2015.
  21. ↑ Grig, Sophie. "Remote Jarawa tribe kill poacher – exclusive interview shows Jarawa denouncing poaching on their land". survivalinternational.org. Survival International. Retrieved 30 May 2015.
  22. 1 2 Anosh Malekar, "The case for a linguisitic survey," Infochange Media, August 1, 2011.
  23. ↑ "Genetic Affinities of the Andaman Islanders, a Vanishing Human Population". Current Biology 13: 86–93. January 2003. doi:10.1016/S0960-9822(02)01336-2. PMID 12546781.
  24. ↑ Jayanta Sarkar (1990), The Jarawa, Anthropological Survey of India, ISBN 81-7046-080-8, The Great Andamanese population was large till 1858 when it started declining ... In 1901, their number was reduced to only 600 and in 1961 to a mere 19.
  25. 1 2 "Traditional handicrafts of Ang tribes (Jarawa) of Andaman Islands" (PDF).
  26. ↑ "A Bibliographical Introduction to Andamanese Linguistics on JSTOR" (PDF). www.jstor.org. Retrieved 2015-10-29.
  27. ↑ Blevins, Juliette (2007). "A Long Lost Sister of Proto-Austronesian?: Proto-Ongan, Mother of Jarawa and Onge of the Andaman Islands.". Oceanic Linguistics 46: 154–198. doi:10.1353/ol.2007.0015.
  28. ↑ "A Grammar of the Great Andamanese Language : An Ethnolinguistic Study". eds.b.ebscohost.com. Retrieved 2015-10-29.
  29. ↑ Man, Edward Horace; Ellis, Alexander John (1932-01-01). The Aboriginal Inhabitants of the Andaman Islands. Mittal Publications.
  30. ↑ "EBSCO Publishing Service Selection Page". eds.b.ebscohost.com. Retrieved 2015-10-30.
  31. 1 2 3 4 5 6 Bulbeck, David (November 2013). "Craniodental Affinities of Southeast Asia's "Negritos" and the Concordance with Their Genetic Affinities". Human Biology 85 (1): 95–134. doi:10.3378/027.085.0305. Retrieved 17 December 2013.
  32. 1 2 "Logging into the proxy: NetID (Rutgers University Libraries)". content.ebscohost.com.proxy.libraries.rutgers.edu. Retrieved 2015-10-29.
  33. 1 2 "Logging into the proxy: NetID (Rutgers University Libraries)". content.ebscohost.com.proxy.libraries.rutgers.edu. Retrieved 2015-10-29.
  34. ↑ "Logging into the proxy: NetID (Rutgers University Libraries)". content.ebscohost.com.proxy.libraries.rutgers.edu. Retrieved 2015-10-29.
  35. 1 2 V. K. Kashyap, Sitalaximi T., B. N. Sarkar, R. Trivedi1 (2003), "Molecular Relatedness of The Aboriginal Groups of Andaman and Nicobar Islands with Similar Ethnic Populations" (PDF), International Journal of Human Genetics 3 (1): 5–11, retrieved 2009-06-08, ... the Negrito populations of Andaman Islands have remained in isolation ... the Andamanese are more closely related to other Asians than to modern day Africans ... the Nicobarese exhibiting a close affinity with geographically proximate Indo-Mongoloid populations of Northeast India ...
  36. 1 2 3 4 M. Phillip Endicott, Thomas P. Gilbert, Chris Stringer, Carles Lalueza-Fox, Eske Willerslev, Anders J. Hansen, Alan Cooper (2003), "The Genetic Origins of the Andaman Islanders" (PDF), American Journal of Human Genetics 72 (1): 178–184, doi:10.1086/345487, PMC 378623, PMID 12478481, retrieved 2009-04-21, ... The HVR-1 data separate them into two lineages, identified on the Indian mainland (Bamshad et al. 2001) as M4 and M2 ... The Andamanese M2 contains two haplotypes ... developed in situ, after an early colonization ... Alternatively, it is possible that the haplotypes have become extinct in India or are present at a low frequency and have not yet been sampled, but, in each case, an early settlement of the Andaman Islands by an M2-bearing population is implied ... The Andaman M4 haplotype ... is still present among populations in India, suggesting it was subject to the late Pleistocene population expansions ...
  37. ↑ Aghakhanian, F.; Yunus, Y.; Naidu, R.; Jinam, T.; Manica, A.; Hoh, B. P.; Phipps, M. E. (14 April 2015). "Unravelling the Genetic History of Negritos and Indigenous Populations of Southeast Asia" (PDF). Genome Biology and Evolution 7 (5): 1206–1215. doi:10.1093/gbe/evv065. Retrieved 1 November 2015.
  38. 1 2 3 4 Reich, David; Kumarasamy Thangaraj; Nick Patterson; Alkes L. Price; Lalji Singh (24 September 2009). "Reconstructing Indian Population History". Nature 461 (7263): 489–494. doi:10.1038/nature08365. PMC 2842210. PMID 19779445. Retrieved 12 November 2013.
  39. ↑ Moorjani, Priya; Kumarasamy Thangaraj; Nick Patterson; Alkes L. Price; Lalji Singh; David Reich (5 September 2013). "Genetic Evidence for Recent Population Mixture in India". American Journal of Human Genetics 93 (3): 422–438. doi:10.1016/j.ajhg.2013.07.006. PMC 3769933. PMID 23932107.
  40. 1 2 "Y-DNA Haplogroup D and its Subclades - 2008". Isogg.org. Retrieved 2010-05-14.
  41. ↑ Wen, Bo; et al. (May 2004). "Analyses of Genetic Structure of Tibeto-Burman Populations Reveals Sex-Biased Admixture in Southern Tibeto-Burmans". American Journal of Human Genetics 5 (74): 856–865. doi:10.1086/386292. PMC 1181980. PMID 15042512. Retrieved 17 December 2013.
  42. ↑ Tajima, Atsushi; et al. (2004), "Genetic origins of the Ainu inferred from combined DNA analyses of maternal and paternal lineages", Journal of Human Genetics 49 (4): 187–193, doi:10.1007/s10038-004-0131-x, PMID 14997363
  43. 1 2 3 Kumarasamy Thangaraj, Lalji Singh, Alla G. Reddy, V. Raghavendra Rao, Subhash C. Sehgal, Peter A. Underhill, Melanie Pierson, Ian G. Frame, and Erika Hagelberg (2002), Genetic Affinities of the Andaman Islanders, a Vanishing Human Population (PDF), archived (PDF) from the original on 29 October 2008, retrieved 2008-11-16, ... Our data indicate that the Andamanese have closer affinities to Asian than to African populations and suggest that they are the descendants of the early Palaeolithic colonizers of Southeast Asia ... All Onge and Jarawa had the same binary haplotype D ... Great Andaman males had five different binary haplotypes, found previously in Southeast Asia, the Indian subcontinent, and Melanesia ...
  44. ↑ Endicott, Phillip; Metspalu, Mait; Stringer, Chris; Macaulay, Vincent; Cooper, Alan; Sanchez, Juan J. (2006-12-20). "Multiplexed SNP Typing of Ancient DNA Clarifies the Origin of Andaman mtDNA Haplogroups amongst South Asian Tribal Populations". PLoS ONE 1 (1): e81. doi:10.1371/journal.pone.0000081. PMC 1766372. PMID 17218991.
  45. ↑ Michael D. Petraglia, Bridget Allchin (2007), The evolution and history of human populations in South Asia, Springer, ISBN 1-4020-5561-7, ... As haplogroup M, except for the African sub-clade M1, is not notably present in regions west of the Indian subcontinent, while it covers the majority of Indian mtDNA variation ...
  46. ↑ Revathi Rajkumar et al., Phylogeny and antiquity of M macrohaplogroup inferred from complete mt DNA sequence of Indian specific lineages, BMC Evolutionary Biology 2005, 5:26 doi:10.1186/1471-2148-5-26 PMID 15804362
  47. ↑ Phillip Endicott, M. Thomas P. Gilbert, Chris Stringer, Carles Lalueza-Fox, Eske Willerslev, Anders J. Hansen, and Alan Cooper (1 January 2002), "The Genetic Origins of the Andaman Islanders", American Journal of Human Genetics (The American Society of Human Genetics) 72 (1): 178–84, doi:10.1086/345487, PMC 378623, PMID 12478481, ... The high frequency of M2 is consistent with its greater age, and its distribution suggests that many of the populations viewed as the autochthons of India because of their cultural inheritance may also be genetic descendants of the early settlers of southern Asia ...
  48. ↑ Choi, Charles (22 September 2011), Now-Extinct Relative Had Sex with Humans Far and Wide, LiveScience

External links

This article is issued from Wikipedia - version of the Monday, April 25, 2016. The text is available under the Creative Commons Attribution/Share Alike but additional terms may apply for the media files.