Leopard attack

The Gunsore man-eater after it was shot by British officer W. A. Conduitt on 21 April 1901. Credited with at least 20 human deaths, the leopard was killed on top of its last victim, a child from Somnapur village in the Seoni district, India.^[1]

Leopard attacks on humans are generally rare occurrences. Despite the leopard's (Panthera pardus) extensive range from sub-Saharan Africa to Southeast Asia, attacks are regularly reported only in India and Nepal.^[2][3] Among the five "big cats", leopards are less likely to become man-eaters—only jaguars and snow leopards have a less fearsome reputation.^[4][5] However, leopards are established predators of non-human primates, sometimes preying on species as large as the western lowland gorilla.^[6] Other primates may make up 80% of the leopard's diet.^[7] While leopards generally avoid humans, they tolerate proximity to humans better than lions and tigers and often come into conflict with humans when raiding livestock.^[8]

Leopard attacks may have peaked in India during the late nineteenth and early twentieth centuries, coinciding with rapid urbanization.^[4] Attacks in India are still relatively common, and in some regions of the country leopards kill more humans than all other large carnivores combined.^[9][10] The Indian states of Gujarat, Himachal Pradesh, Maharashtra, Uttarakhand, and West Bengal experience the most severe human–leopard conflict. In Nepal, most attacks occur in the midland regions (the Terai, midhills, and lesser Himalaya).^[3] One study concluded that the rate of leopard predation on humans in Nepal is 16 times higher than anywhere else, resulting in approximately 1.9 human deaths annually per million inhabitants.^[3] In the former Soviet Central Asia, leopard attacks have been reported in the Caucasus, Turkmenia (present day Turkmenistan), and the Lankaran region of present-day Azerbaijan.^[11] Rare attacks have occurred in China.^[12]

It is possible for humans to win a fight with a leopard, as in the case of a 56-year-old woman who killed an attacking leopard with a sickle and spade, and survived with heavy injuries,^[13] and the case of a 73-year-old man in Kenya who fatally tore the tongue out of a leopard.^[14] Globally, attacks on humans—especially nonfatal attacks that result in only minor injury—likely remain under-reported due to the lack of monitoring programs and standardized reporting protocol.^[15]

Leopard predation on hominids

In 1970, South African paleontologist C. K. Brain showed that a juvenile Paranthropus robustus individual, SK 54, had been killed by a leopard at Swartkrans in Gauteng, South Africa approximately 1.8 million years ago.^[16][17] The SK 54 cranium bears two holes in the back of the skull—holes that perfectly match the width and spacing of lower leopard canine teeth. The leopard appears to have dragged its kill into a tree to eat in seclusion, much like leopards do today.^[16] Numerous leopard fossils have been found at the site, suggesting that the felids were predators of early hominids.^[18] The revelation that these injuries were not the result of interpersonal aggression but were leopard-inflicted dealt a fatal blow to the then-popular killer ape theory.^[19] Another hominid fossil consisting of a 6-million-year-old Orrorin tugenensis femur (BAR 1003'00), recovered from the Tugen Hills in Kenya, preserves puncture damage tentatively identified as leopard bite marks.^[20] This fossil evidence, along with modern studies of primate–leopard interaction, has fueled speculation that leopard predation played a major role in primate evolution, particularly on cognitive development.^[21]

Human–leopard conflict

Reducing human–leopard conflict has proven difficult. Conflict tends to increase during periods of drought or when the leopard's natural prey becomes scarce. Shrinking leopard habitat and growing human populations also increase conflict. In Uganda, retaliatory attacks on humans increased when starving villagers began expropriating leopard kills (a feeding strategy known as kleptoparasitism).^[22] The economic damage resulting from loss of livestock to carnivores caused villagers in Bhutan's Jigme Singye Wangchuck National Park to lose more than two-thirds of their annual cash income in 2000, with leopards blamed for 53% of the losses.^[23] Similarly, in the Annapurna Conservation Area of Nepal, the estimated monetary loss per household was US$95 in 2009 and US$42 in 2010 (out of an annual income of less than US$100), with leopards blamed for 94.9% of the losses.^[24] Like other large carnivores, leopards are capable of surplus killing. Under normal conditions, prey are too scarce for this behavior, but when the opportunity presents itself leopards may instinctually kill in excess for later consumption.^[25] One leopard in Cape Province, South Africa killed 51 sheep and lambs in a single incident.^[26]

Translocation (the capture, transport, and release) of "problem leopards", as with other territorial felids, is generally ineffective: translocated leopards either immediately return or other leopards move in and claim the vacant territory. One translocated leopard in Cape Province traveled nearly 500 kilometres (310 mi) to return to his old territory.^[27] Translocations are also expensive, tend to result in high mortality (up to 70%), and may make leopards more aggressive towards humans, thus failing as both a management and a conservation strategy.^{[28][29][30][31]} Historically, lethal control of problem animals was the primary method of conflict management. Although this remains the situation in many countries,^[30] leopards are afforded the highest legal protection in India under the Wildlife Protection Act of 1972—only man-eaters can be killed and only when they are considered likely to continue to prey on humans.^[32] In Uttarakhand, the state with the most severe human–leopard conflict, 45 leopards were legally declared man-eaters and shot by wildlife officials between 2001 and 2010.^[2]

Where legal, herders may shoot at leopards who prey on their livestock. An injured leopard may become an exclusive predator of livestock if it is unable to kill normal prey, since domesticated animals typically lack natural defenses.^[33] Frequent livestock-raiding may cause leopards to lose their fear of humans, and shooting injuries may have caused some leopards to become man-eaters. There has been increasing acceptance that the "problem leopard" paradigm may be anthropomorphization of normal carnivore behavior, and that translocations are unlikely to stop livestock depredation.^[2][33] In an effort to reduce the shooting of "problem leopards" and lessen the financial burden on herders, some governments provide monetary compensation, although the sum is often less than the value of the lost livestock.^[2]

Number of Human Deaths due to Leopard Attacks^†

Country	Region	Deaths	Year(s)	Ref
India	Indian subcontinent ^‡	11,909	1875–1912	[34]
	Bhagalpur district, Bihar	350	1959–1962	[8]
	Uttarakhand	239	2000–2007	[35]
	Throughout India (mainly Uttarakhand)	170	1982–1989	[36]
	Pauri Garhwal district, Uttarakhand	140	1988–2000	[37]
	Garhwal division, Uttarakhand	125	1918–1926	[38]
	Gujarat	105	1994–2007	[39]
	Uttar Pradesh	95	1988–1998	[19]
	Junagadh district, Gujarat	29	1990–2012	[40]
	Pune district, Maharashtra	18	2001–2003	[10]
	Jammu and Kashmir	17	2004–2007	[41]
	Sanjay Gandhi National Park, Maharashtra	16	1986–1996	[8]
	North Bengal	15	1990–2008	[42]
	Mandi district, Himachal Pradesh	13	1987–2007	[43]
	Chikkamagaluru district, Karnataka	11	1995	[10]
	Kanha National Park, Madhya Pradesh	8	1961–1965	[44]
	Himachal Pradesh	6	2000–2007	[35]
Nepal	Baitadi district, Mahakali zone	15	2010–2012	[45]
	Pokhara Valley, Gandaki zone	12	1987–1989	[46]
Pakistan	Ayubia National Park, Khyber Pakhtunkhwa	12	1989–2006	[47]
	Machiara National Park, Azad Kashmir	2	2004–2007	[48]
Somalia	Golis Mountains, Togdheer	100	c. 1889	[49]
South Africa	Kruger National Park	5	1992–2003	[50]
Sri Lanka	Punanai, Batticaloa district	12	1923–1924	[51]
Zambia	Chambezi River	67	1936–1937	[52]
	Luangwa River	8	1938	[52]
^† No comprehensive global database of fatal leopard attacks exists, and many countries do not keep official records. Due to the fragmentary nature of the data, the deaths reproduced here should be considered minimum figures only. ^‡ The territories forming British India (Bangladesh, Burma, India, and Pakistan)

Man-eaters

Characteristics

The leopard is largely a nocturnal hunter. For its size, it is the most powerful large felid, able to drag a carcass larger than itself up a tree.^[53] Leopards can run more than 60 kilometres per hour (37 mph), leap more than 6 metres (20 ft) horizontally and 3 metres (9.8 ft) vertically, and have a more developed sense of smell than tigers.^[53] They are strong climbers and can descend down a tree headfirst.^[53] Man-eating leopards have earned a reputation as being particularly bold and difficult to track. British hunters Jim Corbett (1875–1955) and Kenneth Anderson (1910–1974) wrote that hunting leopards presented more challenges than any other animal.^[54][55] Indian naturalist J. C. Daniel (1927–2011), former curator of the Bombay Natural History Society, reprinted many early twentieth-century accounts of man-eating leopards in his book The Leopard in India: A Natural History (Dehradun: Natraj Publishers, 2009). One such account in the Journal of the Bombay Natural History Society describes the unique danger posed by leopards:

Like the tiger, the panther [leopard] sometimes takes to man-eating, and a man-eating panther is even more to be dreaded than a tiger with similar tastes, on account of its greater agility, and also its greater stealthiness and silence. It can stalk and jump, and...can climb better than a tiger, and it can also conceal itself in astonishingly meager cover, often displaying uncanny intelligence in this act. A man-eating panther frequently breaks through the frail walls of village huts and carries away children and even adults as they lie asleep.^[8]

One study concluded that only 9 of 152 documented man-eating leopards were female.^[56] Drawing on the sex and physical condition of 78 man-eating leopards, the same study concluded that man-eaters were typically uninjured mature males (79.5%), with a fewer number of aged and immature males (11.6% and 3.8%, respectively).^[56] Once a leopard has killed and eaten a human, they are likely to persist as man-eaters—they may even show a nearly exclusive preference for humans.^[17] In "Man-Eaters of Kumaon", Jim Corbett mentioned that leopards are driven to man-eating by acquiring a taste for human flesh due to scavenging on corpses thrown into the jungle during an epidemic. He wrote,"A leopard, in an area in which his natural food is scarce, finding these bodies very soon acquires a taste for human flesh, and when the disease dies down and normal conditions are established, he very naturally, on finding his food supply cut off, takes to killing human beings". Of the two man-eating leopards of Kumaon, which between them killed five hundred and twenty-five people, the Panar Leopard followed on the heels of a very severe outbreak of cholera, while the Rudraprayag Leopard followed the 1918 influenza epidemic which was particularly deadly in India.^[57] Corbett wrote that the Rudraprayag man-eater once broke into a pen holding 40 goats, but instead of attacking the livestock it killed and ate the sleeping 14-year-old boy who had been assigned to guard them.

Leopard attacks on humans tend to occur at night, and often close to villages. There have been documented incidents of leopards forcing their way into human dwellings at night and attacking the inhabitants in their sleep.^[15] A number of fatal attacks have also occurred in zoos and homes with pet leopards.^{[58][59][60][61]} During predatory attacks, leopards typically bite their prey's throat or the nape of the neck, lacerating or severing jugular veins and carotid arteries, causing rapid exsanguination. The spine may be crushed and the skull perforated, exposing the brain.^[41][58][59] Survivors of attacks typically suffer extensive trauma to the head, neck, and face. Multibacterial infection resulting from the contamination of wounds by leopard oral flora occurs in 5–30% of attack survivors, complicating recovery.^[58] Before the advent of antibiotics, 75% of attack survivors died from infection.^[62]

Notable man-eaters

The Panar Leopard killed by Jim Corbett

British hunter Jim Corbett poses after shooting the Rudraprayag leopard on 2 May 1926

• Leopard of Panar : The Leopard of Panar was a male leopard reported as being responsible for at least 400 fatal attacks on humans in the Panar region of the Almora district, situated in Kumaon Northern India in the early 20th century. Jim Corbett heard of the leopard while hunting the Champawat tiger in 1907, and in 1910 he set out to kill it. Although it apparently claimed hundreds of more lives than the Rudraprayag man-eater, the Panar man-eater received less attention from the British Indian press, which Corbett attributed to the remoteness of Almora.^[63]
• Leopard of the Central Provinces
• Leopard of Rudraprayag
• Leopard of Gummalapur
• Leopard of the Yellagiri Hills
• Leopard of the Golis Range: In 1899 British officer H. G. C. Swayne (1860–1940) wrote of a man-eating leopard that had allegedly killed more than 100 humans in the Golis Mountains of British Somaliland. Swayne's brief account appears in the volume Great and Small Game of Africa (London: Roland Ward, 1899), edited by the prominent British naturalist Henry Bryden (1854–1937):

  In 1889 there was a leopard, said to be a panther, which had haunted the Mirso ledge of the Golis range for some years, and was supposed to have killed over a hundred people. It was in the habit of lying in wait at a corner of a very dark, rough jungle path, where huge rocks overlooked the track; and the Somalis used to show a boulder, some 6 feet high, a yard from the path, in the flat top of which was a depression shaped like a panther's body, from which the beast was said to spring upon travellers.^[49]

  According to Swayne, leopards were more abundant in the Golis Mountains than anywhere else in British Somaliland, and were responsible for 90% of all attacks on sheep and goats. The rocky terrain of the Golis made tracking and killing leopards next to impossible.^[49] At the time of the attacks, this remote territory remained largely unexplored by the British, and little else is known of the Golis Range man-eater.
• Leopard of the Mulher Valley: In 1903 L. S. Osmaston (1870–1969), a conservator employed by the Imperial Forestry Service, reported that a man-eating leopard had killed more than 30 humans in the Mulher Valley between 1901 and 1902.^[64] Osmaston twice set out to kill the leopard in February and March 1902, but was unsuccessful. His forestry work required him to leave Mulher later that month, and he was unable to return until late November. The leopard's last attack occurred a few days later on 3 December:

  I heard a boy of 15 had been killed at Wadai, 4 miles from my camp; this boy was most unfortunate. Last year the panther had tried to get him, but only mauled one leg; my wife and I were able to dose the wound with carbolic oil and the boy got well; this time he and one or two others were sitting close to a bright fire on a threshing floor near the village in the early part of the night and the panther came and carried him off: the panther took him about a quarter of a mile to a patch of high grass and brushwood and ate all he could of the head, the flesh of one leg and all his inside; so there was plenty left for the beast to come back for.^[64]

  Osmaston constructed a blind 11 metres (36 ft) from the boy's corpse and waited. The leopard returned to the area in the afternoon, but cautiously avoided approaching the body until after dark. When it finally ventured within shooting range, Osmaston fired with his double-barreled express rifle. The injured animal darted off into the night, and was killed the following morning when it was discovered alive some distance away. Osmaston speculated that the attacks began during the Indian famine of 1899–1900, the leopard having taken to man-eating after killing a dying person in the jungle. He also believed the man-eater was responsible for other fatal attacks in the nearby Dang and Dhule districts, but did not know the exact number of fatalities.^[64]
• Leopard of Kahani: Robert A. Sterndale (1839–1902) and James Forsyth (1838–1871) gave accounts of a man-eating leopard that killed "nearly a hundred persons" in the Seoni district between 1857 and 1860. When Sterndale received word of the attacks he pursued the man-eater with his brother-in-law, W. Brooke Thomson, but their efforts proved fruitless.^[65] The breakout of the Indian Rebellion of 1857 sent Sterndale away for two years and ended his chance to capture the man-eater. The leopard evaded all attempts by locals to kill it and terrorized the villages of Dhuma and Kahani, sometimes killing three humans in a single night.^[65] According to Sterndale, the leopard preferred to consume blood rather than flesh, and most bodies showed few injuries other than telltale bite marks to the throat.^[65] A large reward was offered for the leopard's capture, and it was then unexpectedly killed one night by an inexperienced native hunter.^[65] When Forsyth passed through Seoni several years later, the leopard's story had become legendary. He later recounted a myth he had heard from the locals:

  A man and his wife were travelling back to their home from a pilgrimage to Benares, when they met on the road a panther. The woman was terrified; but the man said, "Fear not, I possess a charm by which I can transform myself into any shape. I will now become a panther, and remove this obstacle from the road, and on my return you must place this powder in my mouth, when I will recover my proper shape." He then swallowed his own portion of the magic powder, and assuming the likeness of the panther, persuaded him to leave the path. Returning to the woman, he opened his mouth to receive the transposing charm; but she, terrified by his dreadful appearance and open jaws, dropped it in the mire, and it was lost. Then, in despair, he killed the author of his misfortune, and ever after revenged himself on the race whose form he could never resume.^[66]

Recent attacks

On January 6, 2015, a leopard injured a boy in Katarniaghat Wildlife Sanctuary, Uttar Pradesh.^[67] Another leopard killed 2 men in Kolar district, Karnataka.^[68] On January 13, 2015, a leopard injured a woman with paw in Jiuni Valley of Mandi district while she had gone to collect fodder for household animals, injuring her on the neck.^[69] On January 16, 2015, 2 boys were killed by a leopard in Uttarakhand.^[70] On January 25, 2015, a leopard killed a girl in Galyat.^[71] The next day, a leopard mauled 5 people, seriously injuring 3, before being killed in Jalpaiguri district.^[72]

On February 14, 2015, a leopard injured 2 villagers in Sagar district.^[73] Four days later, a leopard injured 6 people in Shravasti district.^[74] On February 22, 2015, a girl was injured on the head when a leopard attacked her in Dingore village (approximately 20 km from Junnar).^[75] On March 5, 2015, a leopardess critically injured 4 people before being stabbed to death in a forest of Jorhat district.^[76]

See also

• Human–wildlife conflict
• Anthrozoology

References

1. Conduitt, W. A. (1903). "A man-eating panther". Journal of the Bombay Natural History Society 14: 595–597. Retrieved 22 March 2013.
2. Athreya, V. (2012). Conflict resolution and leopard conservation in a human dominated landscape” (Ph.D.). Manipal University. Retrieved 29 March 2013. C1 control character in |title= at position 76 (help)
3. Maskey, T. M.; Bauer, J.; Cosgriff, K. (2001). Village children, leopards and conservation. Patterns of loss of human live through leopards (Panthera pardus) in Nepal (Report). Kathmandu, Nepal: Department of National Parks and Wildlife Conservation/Sustainable Tourism CRC.
4. Quigley, H.; Herrero, S. (2005). "Chapter 3: Characterization and prevention of attacks on humans". In Woodroffe, R.; Thirgood, S.; Rabinowitz, A. People and wildlife: Conflict or co-existence?. Cambridge University Press. pp. 27–48. ISBN 9780521825054.
5. Inskip, C.; Zimmermann, A. (2009). "Human-felid conflict: A review of patterns and priorities worldwide". Oryx 43 (1): 18–34. doi:10.1017/S003060530899030X.
6. Fay, J. M.; Carroll, R.; Kerbis-Peterhans, J. C.; Harris, D. (1995). "Leopard attack on and consumption of gorillas in the Central African Republic". Journal of Human Evolution 29 (1): 93–99. doi:10.1006/jhev.1995.1048.
7. Srivastava, K. K.; Bhardwaj, A. K.; Abraham, C. J.; Zacharias, V. J. (1996). "Food habits of mammalian predators in Periyar Tiger Reserve, South India". The Indian Forester 122 (10): 877–883. Retrieved 22 March 2013.
8. Quammen, D. (2003). Monster of God: The man-eating predator in the jungles of history and the mind. New York: W. W. Norton & Company. pp. 55–61. ISBN 9780393326093. Retrieved 20 March 2013.
9. Kimothi, P. (February 5, 2011). "Losers on both sides as man-animal war rages". The Pioneer. Archived from the original on March 2, 2011. Retrieved 22 March 2013.
10. Athreya, V. R.; Thakur, S. S.; Chaudhuri, S.; Belsare, A. V. (2004). A study of the man-leopard conflict in the Junnar Forest Division, Pune District, Maharashtra (PDF) (Report). Submitted to the Office of the Chief Wildlife Warden, Maharashtra State Forest Department, and the Wildlife Protection Society of India, New Delhi, India. Retrieved 22 March 2013.
11. Heptner, V.G.; and Sluskii, A.A. Mammals of the Soviet Union. Vol 2, Part 2. (Carnivores: Hyaenas and Cats) New Delhi: Amerind Publishing; 1992. p269-271.
12. Heptner, V.G.; and Sluskii, A.A. Mammals of the Soviet Union. Vol 2, Part 2. (Carnivores: Hyaenas and Cats) New Delhi: Amerind Publishing; 1992. p269.
13. "Indian woman survives leopard attack after fighting predator for 30 minutes". The Telegraph. 27 August 2014. Retrieved 27 August 2014.
14. http://www.news24.com/Africa/News/Man-rips-leopards-tongue-out-20050622
15. Löe, J.; Röskaft, E. (2004). "Large carnivores and human safety: A review". AMBIO: A Journal of the Human Environment 33 (6): 283–288. doi:10.1579/0044-7447-33.6.283.
16. Brain, C. K. (1970). "New finds at the Swartkrans australopithecine site". Nature 225 (5238): 1112–1119. doi:10.1038/2251112a0.
17. Brain, C. K. (1981). The hunters or the hunted?: An introduction to African cave taphonomy. Chicago: University of Chicago Press. pp. 97–98, 266–274. ISBN 9780226070896.
18. Lee-Thorp, J.; Thackeray, J. F.; Van der Merwe, N. (2000). "The hunters and the hunted revisited". Journal of Human Evolution 39 (6): 565–576. doi:10.1006/jhev.2000.0436.
19. Hart, D. L.; Sussman, R. W. (2005). Man the hunted: Primates, predators, and human evolution. Cambridge, Massachusetts: Westview Press. pp. 1–11, 60–62. ISBN 9780813339368.
20. Gommery, G.; Pickford, M.; Senut, B. (2007). "A case of carnivore-inflicted damage to a fossil femur from Swartkrans, comparable to that on a hominid femur representing Orrorin tugenensis, BAR 1003'00 (Kenya)". Annals of the Transvaal Museum 44: 215–218. ISSN 0041-1752. Retrieved 22 March 2013.
21. Zuberbühler, K.; Jenny, D. (2002). "Leopard predation and primate evolution". Journal of Human Evolution 43 (6): 873–886. doi:10.1006/jhev.2002.0605.
22. Treves, A.; Naughton-Treves, L. (2009). "Risk and opportunity for humans coexisting with large carnivores". Journal of Human Evolution 36 (3): 275–282. doi:10.1006/jhev.1998.0268.
23. Wang, S. W.; Macdonald, D. W. (2006). "Livestock predation by carnivores in Jigme Singye Wangchuck National Park, Bhutan". Biological Conservation 129 (4): 558–565. doi:10.1016/j.biocon.2005.11.024.
24. Koirala, R. K.; Aryal, A.; Parajuli, A.; Raubenheimer, D. (2012). "Human-common leopard (Panthera pardus) conflict in lower belt of Annapurna Conservation Area, Nepal" (PDF). Journal of Research in Conservation Biology (Ficus Publishers) 1 (1): 5–12. Retrieved 29 March 2013.
25. Kruuk, H. (2002). Hunter and hunted: Relationships between carnivores and people. Cambridge: Cambridge University Press. pp. 51–53, 58–60, 104. ISBN 9780521814102.
26. Stuart, C. T. (1986). "The incidence of surplus killing by Panthera pardus and Felis caracal in Cape Province, South Africa". Mammalia 50 (4): 556–558. doi:10.1515/mamm.1986.50.4.553. ISSN 0025-1461.
27. Jewell, P. A. (1982). Conservation of the cheetah: Should cheetah be moved to distant areas? (PDF) (Report). Cambridge: Unpublished workshop report, International Fund for Animal Welfare. Retrieved 27 March 2013.
28. Athreya, V.; Odden, M.; Linnell, J. D. C.; Karanth, K. U. (2011). "Translocation as a tool for mitigating conflict with leopards in human-dominated landscapes of India". Conservation Biology 25 (1): 133–141. doi:10.1111/j.1523-1739.2010.01599.x.
29. Athreya, V. R.; Thakur, S. S.; Chaudhuri, S.; Belsare, A. V. (2007). "Leopards in human-dominated areas: A spillover from sustained translocations into nearby forests?". Journal of the Bombay Natural History Society 104 (1): 45–50. ISSN 0006-6982.
30. Treves, A.; Karanth, K. U. (2003). "Human-carnivore conflict and perspectives on carnivore management worldwide". Conservation Biology 17 (6): 1491–1499. doi:10.1111/j.1523-1739.2003.00059.x.
31. Linnell, J. D. C.; Aanes, R.; Swenson, J. E.; Odden, J.; Smith, M. E. (1997). "Translocation of carnivores as a method for managing problem animals: A review". Biodiversity and Conservation 6 (9): 1245–1257. doi:10.1023/B:BIOC.0000034011.05412.cd.
32. Guidelines for human-leopard conflict management (PDF) (Report). Ministry of Environment and Forests, Government of India. 2011. p. 17. Retrieved 30 March 2013.
33. Linnell, J. D. C.; Odden, J.; Smith, M. E.; Aanes, R.; Swenson, J. E. (1999). "Large carnivores that kill livestock: Do "problem individuals" really exist?". Wildlife Society Bulletin 27 (3): 698–705. Retrieved 29 March 2013.
34. Compiled from official British records available at the Digital South Asia Library (University of Chicago and the Center for Research Libraries).
    1. "Number of persons and cattle killed in British India by wild beasts and snakes", Statistical abstract relating to British India from 1867–68 to 1876–77, (London: Her Majesty's Stationery Office): p. 132, 1878, retrieved 30 March 2013.
    2. "Number of persons and cattle killed in British India by wild beasts and snakes", Statistical abstract relating to British India from 1876–77 to 1885–86, (London: Her Majesty's Stationery Office): p. 240, 1887, retrieved 30 March 2013.
    3. "Number of persons and cattle killed in British India by wild beasts and snakes", Statistical abstract relating to British India from 1885–86 to 1894–95, (London: Her Majesty's Stationery Office): p. 268, 1896, retrieved 30 March 2013.
    4. "Number of persons and cattle killed in British India by wild animals and snakes", Statistical abstract relating to British India from 1894–95 to 1903–04, (London: Her Majesty's Stationery Office): p. 238, 1905, retrieved 30 March 2013.
    5. "Number of persons and cattle killed in British India by wild animals and snakes", Statistical abstract relating to British India from 1903–04 to 1912–13, (London: His Majesty's Stationery Office): p. 240, 1915, retrieved 30 March 2013.
35. Marker, L.; Sivamani, S. (2009). "Policy for human-leopard conflict management in India" (PDF). Cat News 50: 23–26. Retrieved 22 March 2013.
36. Johnsingh, A. J. T.; Panwar, H. S.; Rodgers, W. A. (1991). "Ecology and conservation of large felids in India". In Maruyama, N.; Bobek, B.; Ono, Y.; Regelin, W.; Bartos, L.; Ratcliffe, P. R. Wildlife conservation: Present trends and perspectives for the 21st century. Yokohama: Japan Wildlife Research Center. pp. 160–165. OCLC 749891670.
37. Goyal, S. P.; Chauhan, D. S.; Agrawal, M. K.; Thapa, R. (2000). A study on distribution, relative abundance and food habits of leopard (Panthera pardus) in Garhwal Himalayas (PDF) (Report). Dehradun: Wildlife Institute of India. Retrieved 22 March 2013.
38. Corbett, E. J. (1948). The man-eating leopard of Rudraprayag. London: Oxford University Press. OCLC 424546. Retrieved 29 March 2013.
39. 1. "Man–animal conflict". Forest Department, Government of Gujarat. 20 July 2012. Archived from the original on July 23, 2012. Retrieved 29 March 2013.
    2. Marker, L.; Sivamani, S. (2009). "Policy for human-leopard conflict management in India" (PDF). Cat News 50: 23–26. Retrieved 22 March 2013.
40. 1. Shastri, P. (February 25, 2013). "Leopards kill 12 in Junagadh, injure 48 in one year". The Times of India. Retrieved 21 March 2013.
    2. Vijayan, S.; Pati, B. P. (2002). "Impact of changing cropping patterns on man–animal conflicts around Gir Protected Area with specific reference to Talala sub-district, Gujarat, India". Population and Environment 23 (6): 541–559. doi:10.1023/A:1016317819552.
41. Nabi, D. G.; Tak, S. R.; Kangoo, K. A.; Halwai, M. A. (2009). "Injuries from leopard attacks in Kashmir". Injury 40 (1): 90–92. doi:10.1016/j.injury.2008.05.033.
42. 1. WWF–India (1997). Leopard study report (Report). New Delhi, India: World Wide Fund for Nature. p. 49.
    2. Bhattacharjee, A.; Parthasarathy, N. (2013). "Coexisting with large carnivores: A case study from western Duars, India". Human Dimensions of Wildlife 18 (1): 20–31. doi:10.1080/10871209.2012.698403.
43. Kumar, D.; Chauhan, N. P. S. (2011). "Human-leopard conflict in Mandi district, Himachal Pradesh, India". Julius-Kühn-Archiv 432: 180–181. doi:10.5073/jka.2011.432.098.
44. Schaller, G. B. (1967). The deer and the tiger: A study of wildlife in India. Chicago: University of Chicago Press. p. 311. ISBN 9780226736310. Retrieved 29 March 2013.
45. Shrestha, M. (November 3, 2012). "Leopard suspected of eating 15 people in Nepal". CNN. Retrieved 21 March 2013.
46. 1. "Leopard attacks in Nepal". Cat News (International Union for Conservation of Nature) 9. 1989. Archived from the original on March 21, 2005. Retrieved 29 March 2013.
    2. "Man-eating leopard in Nepal". Cat News (International Union for Conservation of Nature) 6. 1987. Archived from the original on March 21, 2005. Retrieved 29 March 2013.
    3. "Leopard stoned to death". Cat News (International Union for Conservation of Nature) 11. 1989. Archived from the original on March 18, 2005. Retrieved 29 March 2013.
47. Lodhi, A. (2007). Conservation of leopard in Ayubia National Park, Pakistan (PDF) (M. S.). University of Montana. pp. 19–20. Retrieved 29 March 2013.
48. Dar, N. I.; Minhas, R. A.; Zaman, Q.; Linkie, M. (2009). "Predicting the patterns, perceptions and causes of human–carnivore conflict in and around Machiara National Park, Pakistan". Biological Conservation 142 (10): 2076–2082. doi:10.1016/j.biocon.2009.04.003.
49. Swayne, H. G. C. (1899). "The leopard in Somaliland". In Bryden, H. A. Great and small game of Africa: An account of the distribution, habits, and natural history of the sporting mammals, with personal hunting experiences. London: Roland Ward. pp. 575–579. OCLC 11014130.
50. 1. "Ranger hands it to leopard". News24. 19 December 2003. Retrieved 29 March 2013.
    2. Lubisi, D. (8 March 2001). "Leopard in territory battle?". News24. Retrieved 29 March 2013.
51. Jayewardene, R.; Kumara, J.; Miththapala, S.; Perera, H.; Samarasinha, R.; Santiapillai, C.; Seidensticker, J. (2002). For the leopard: A tribute to the Sri Lankan leopard. Colombo, Sri Lanka: Leopard Trust. p. 30. ISBN 9789558798003.
52. Brelsford, V. (1950). "Unusual Events in Animal Life—IV". African Wild Life 4 (1): 67.
53. Nowak, R. M. (2005). Walker's carnivores of the world. Baltimore, Maryland: Johns Hopkins University Press. pp. 264–268. ISBN 9780801880339.
54. Nowell, K.; Jackson, P., eds. (1996). Wild cats: Status survey and conservation action plan. Gland, Switzerland: International Union for Conservation of Nature. pp. 193–195. ISBN 9782831700458. Retrieved 21 March 2013.
55. Anderson, K. (1955). Nine man-eaters and one rogue. New York: E. P. Dutton. pp. 36–51. OCLC 529362. Retrieved 21 March 2013.
56. Turnbull-Kemp, P. (1967). The leopard. Cape Town: Howard Timmins. pp. 130–146. OCLC 715383208.
57. The Man-eating Leopard of Rudraprayag, Jim Corbett, Oxford university press, ISBN 0-19-562256-1
58. Bahram, R.; Burke, J. E.; Lanzi, G. L. (2004). "Head and neck injury from a leopard attack: Case report and review of the literature". Journal of Oral and Maxillofacial Surgery 62 (2): 247–249. doi:10.1016/j.joms.2003.04.015.
59. Cohle, S. D.; Harlan, C. W.; Harlan, G. (1990). "Fatal big cat attacks". American Journal of Forensic Medicine and Pathology 11 (3): 208–212. doi:10.1097/00000433-199009000-00007. PMID 2220706.
60. Hejna, P. (2010). "A fatal leopard attack". Journal of Forensic Sciences 55 (3): 832–834. doi:10.1111/j.1556-4029.2010.01329.x.
61. Vogel, J. S.; Parker, J. R.; Jordan, F. B.; Coury, T. L.; Vernino, A. R. (2000). "Persian leopard (Panthera pardus) attack in Oklahoma: Case report". American Journal of Forensic Medicine and Pathology 21 (3): 264–269. doi:10.1097/00000433-200009000-00017. PMID 10990290.
62. Auerbach, P. S. (2012). Wilderness Medicine (6th ed.). Philadelphia: Mosby. pp. 1114–1115. ISBN 9781437716788. Retrieved 29 March 2013.
63. Corbett, E. J. (1954). The temple tiger and more man-eaters of Kumaon. London: Oxford University Press. pp. 64–86. OCLC 1862625. Retrieved 29 March 2013.
64. Osmaston, L. S. (1904). "A man-eating panther". Journal of the Bombay Natural History Society 15: 135–138. Retrieved 22 March 2013.
65. Sterndale, R. A. (1877). Seonee; Or, camp life on the Satpura Range: A tale of Indian adventure (2nd ed.). London: Sampson Low, Marston, Searle, & Rivington. pp. 370–384, 452. OCLC 27112858. Retrieved 22 March 2013.
66. Forsyth, J. (1889). The highlands of central India: Notes on their forests and wild tribes, natural history, and sports (New ed.). London: Chapman and Hall. pp. 334–335. OCLC 575941. Retrieved 22 March 2013.
67. http://timesofindia.indiatimes.com/Boy-injured-in-leopard-attack/articleshow/45799926.cms
68. http://www.deccanherald.com/content/452231/man-grandson-die-leopard-attack.html
69. http://www.divyahimachal.com/himachal/sundernagar-mandi-news/%e0%a4%aa%e0%a4%a4%e0%a5%8d%e0%a4%a4%e0%a4%bf%e0%a4%af%e0%a4%be%e0%a4%82-%e0%a4%9c%e0%a5%81%e0%a4%9f%e0%a4%be-%e0%a4%b0%e0%a4%b9%e0%a5%80-%e0%a4%ae%e0%a4%b9%e0%a4%bf%e0%a4%b2%e0%a4%be-%e0%a4%aa/
70. http://www.dailymail.co.uk/indiahome/indianews/article-2914847/Two-boys-aged-3-10-killed-fresh-leopard-attacks-Uttarakhand.html
71. http://tribune.com.pk/story/827523/searching-for-food-girl-killed-in-leopard-attack/
72. http://zeenews.india.com/news/sci-tech/leopard-mauls-five-beaten-to-death_1536834.html
73. http://www.hindustantimes.com/bhopal/two-villagers-injured-in-leopard-attack-in-madhya-pradesh/article1-1316933.aspx
74. http://timesofindia.indiatimes.com/city/allahabad/Several-injured-in-leopard-attacks/articleshow/46293078.cms
75. http://timesofindia.indiatimes.com/city/pune/14-year-old-girl-injured-in-leopard-attack-near-Junnar/articleshow/46361560.cms
76. http://timesofindia.indiatimes.com/city/guwahati/Leopard-beaten-to-death-in-Jorhat/articleshow/46479152.cms