Mesolenellus

Mesolenellus
Temporal range: 516–513 Ma

Botomian

Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Trilobita
Order: Redlichiida
Suborder: Olenellina
Superfamily: Olenelloidea
Family: Olenellidae
Subfamily: Mesonacinae
Genus: Mesolenellus
Palmer & Repina, 1993
Species
  • Mesolenellus hyperborea
    (Poulsen, 1974) type species, synonyms Olenellus hyperborea, Holmia hyperborea[1]
  • Mesolenellus svalbardensis
    (Kielan, 1960), synonym Olenellus svalbardensis[1][2]

Mesolenellus is an extinct genus of trilobites that lived during the lower Cambrian (Botomian), found in Greenland and Spitsbergen.[1]

Etymology

Mesolenellus is a contraction of meso - middle - and Olenellus, the genus from which it was split-off. The epithet hyperborea is a contraction of hyper (Greek ὑπέρ "excess") and Boreas (Greek: Βορέας), god of the North Wind in Greek mythology, referencing the current geographical position of the deposits of its type location. M. svalbardensis has been named for the Norwegian island group Svalbard or Spitsbergen, where it was originally collected.

Taxonomy

It was considered a subgenus of Olenellus before.[3] The nearest relatives of Mesolenellus are the species of the genus Mesonacis, with which it constitutes the subfamily Mesonacinae.

Distribution

Habitat

The mudstone in which both species were found indicates deeper, quiet waters.[5]

Ecology

Mesolenellus hyperborea occurs together with Limniphacos perspiculum, Serrodiscus, hyoliths, Petrianna fulmenta (Bradoriida), and inarticulate brachiopods.[5]

Description

As with most early trilobites, Mesolenellus has an almost flat exoskeleton, that is only thinly calcified, and has crescent-shaped eye ridges. As part of the Olenellina suborder, Olenellus lacks dorsal sutures. Like all other members of the Olenelloidea superfamily, the eye-ridges spring from the back of the frontal lobe (L4) of the central area of the cephalon, that is called glabella. Mesolenellus also shares the typical character of whole Olenellidae family that the frontal (L3) and middle pair (L2) of lateral lobes of the glabella are partially merged. This creates two very typical, isolated slits. Mesolenellus is a genus within the Mesonacinae, with eye-ridges curved but almost parallel to the midline. The back of the eye-ridges is opposite the most backward ring of the glabella (L0 or occipetal ring). Genal spines are 6-8 times as long as L0. The outer furrows of the glabella are parallel to the midline between the back of the cephalon and the furrow between side lobes L2 and L3. The thorax is 3 times as wide as the axis at the 3rd segment. Mesolenellus can be distinguished from the sister-genus Mesonacis, where the back of the eye-ridge extends only to the most backward side lobes (L1), genal spines are only 1-5 times as long as L0, and the glabella widens forwards along L1 and L2. Except for in Mesonacis fremonti the curved eye-ridges are at an angle of 15°-20° with the midline. A third genus Olenellus, that constitutes the second subfamily of the family Olenellidae differs from Mesolenellus in having genal spines 4-5 times as long as L0, the glabella widens forwards along L1 and L2, and the thorax 4-4½ times wider that the axis at the 3rd segment.[1]

Key to the species

1 A short ridge (called plectrum) connects the frontal lobe of the glabella with the frontal border of the headshield. The section of the border of the back of the head (posterior cephalic border) halfway between the midline and the genal angle (between the axis and the intergenal angle) has a backward angle. The base of the genal spine is opposite the most backward ring of the glabella (L0). The furrow between the 2nd and 3rd ring (S2) does not contact the furrow that outlines the glabella (or axial furrow).[1] Nevadella-zone. Svalbard (Vestspitsbergen, Hornsund) and Greenland (Peary Land, Schley Fjord Formation and Buen Formation).
M. svalbardensis (Kielan, 1960)
- Plectrum absent. The midsection of the posterior cephalic border is transverse. The base of the genal spine is opposite the middle ring of the glabella (L2). S2 contacts the axial furrow.[1] Nevadella-zone. Greenland (Buen Formation).
M. hyperborea (Poulsen, 1974)

References

  1. 1 2 3 4 5 6 Lieberman, B.S. (1999). "Systematic Revision of the Olenelloidea (Trilobita, Cambrian)" (PDF). Bulletin of the Peabody Museum of Natural History 45.
  2. Cowie, J.; McNamara, K.J. (1978). "Olenellus (Trilobita) from the Lower Cambrian Strata of North-West Scotland" (PDF). Palaeontology 21 (3): 615–634.
  3. H. B. Whittington; et al. (1997). "Introduction, Order Agnostida, Order Redlichiida". Treatise on Invertebrate Paleontology. Part O, Revised. Trilobita.
  4. Poulsen, V. (1974). "Olenellacean trilobites from eastern North Greenland". Bulletin of the Geological Society of Denmark 23: 79–101.
  5. 1 2 3 4 Mark R. Blaker & John S. Peel (1997). Lower Cambrian trilobites from North Greenland. Meddelelser om Grønland. Geoscience 35. ISBN 978-87-635-1241-1.
  6. Kielan, Z. (1960). "On two olenellid trilobites from Hornsund, Vestspitsvergen". Studia Geologica Polonica 4: 133–139.

External links

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