Pammene

Pammene
Adult Pammene aurana at Commanster (Belgian Ardennes)
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Lepidoptera
Division:	Ditrysia
Family:	Tortricidae
Subfamily:	Olethreutinae
Tribe:	Grapholitini
Genus:	Pammene Hübner, [1825]
Type species
Tortrix trauniana Denis & Schiffermüller, 1775
Diversity
89 species
Synonyms
Numerous, see text

The tortrix moth genus Pammene belongs to the tribe Grapholitini of subfamily Olethreutinae.^[1]

Species

89 species are currently recognized in Pammene:^[2]

Pammene aceris Kuznetzov in Danilevsky & Kuznetsov, 1968 Pammene adusta Kuznetzov 1972 Pammene agnotana Rebel, 1914 Pammene ainorum Kuznetzov in Danilevsky & Kuznetsov, 1968 Pammene albuginana (Guenée, 1845) Pammene amygdalana (Duponchel in Godart, 1842) Pammene argyrana (Hübner, [1799]) Pammene aurana Pammene aurita Razowski, 1992 Pammene avetianae Kuznetzov, 1964 Pammene bathysema Diakonoff, 1976 Pammene blanda Bae & Park, 1998 Pammene blockiana (Herrich-Schäffer, 1851) Pammene bowmanana (McDunnough, 1927) Pammene caeruleata Kuznetzov, 1970 Pammene caliginosa Kuznetzov, 1972 Pammene christophana (Moschler, 1862) Pammene clanculana (Tengström, 1869) Pammene cocciferana Walsingham, 1903 Pammene crataegicola Liu & Komai, 1993 Pammene crataegophila Amsel, 1935 Pammene cyanatra (Diakonoff, 1976) Pammene cytisana (Zeller, 1847) Pammene engadinensis Müller-Rutz, 1920 Pammene epanthista (Meyrick, 1922) Pammene exscribana Kuznetzov in Ler, 1986 Pammene fasciana – Chestnut Leafroller Pammene felicitana Heinrich, 1923 Pammene festiva Kuznetzov, 1972 Pammene flavicellula Kuznetzov, 1971 Pammene fulminea (Komai, 1999) Pammene gallicana (Guenée, 1845) Pammene gallicolana (Lienig & Zeller, 1846) Pammene germmana (Hübner, [1799]) Pammene giganteana (Peyerimhoff, 1863) Pammene ginkgoicola Liu, 1992 Pammene griseana Walsingham, 1900 Pammene griseomaculana Kuznetzov, 1960 Pammene grunini (Kuznetzov, 1960) Pammene ignorata Pammene insolentana Kuznetzov, 1964 Pammene instructana Kuznetzov, 1964 Pammene insulana (Guenée, 1845) Pammene japonica Kuznetzov in Danilevsky & Kuznetsov, 1968 Pammene juniperana (Millière, 1858)

Pammene laserpitiana Huemer & Erlebach, 1999 Pammene leucitis (Meyrick, 1907) Pammene luculentana Kuznetzov in Danilevsky, Kuznetsov & Falkovitsh, 1962 Pammene luedersiana (Sorhagen, 1885) Pammene macrolepis Diakonoff, 1976 Pammene mariana (Zerny, 1920) Pammene medioalbana Knudson, 1986 Pammene megalocephala Diakonoff, 1983 Pammene monotincta Kuznetzov, 1976 Pammene nannodes Walsingham, 1900 Pammene nemorosa Kuznetzov in Danilevsky & Kuznetsov, 1968 Pammene nequior Kuznetzov, 1964 Pammene nescia Kuznetzov, 1972 Pammene nigritana (Mann, 1862) Pammene obscurana (Stephens, 1834) Pammene ochsenheimeriana (Lienig & Zeller, 1846) Pammene ocliferia (Heinrich, 1926) Pammene oreina Kuznetzov, 1973 Pammene orientana Kuznetzov, 1960 Pammene ornata Walsingham, 1903 Pammene oxycedrana (Millière, 1876) Pammene paula (Heinrich, 1926) Pammene percognata Diakonoff, 1976 Pammene perstructana (Walker, 1863) Pammene phthoneris Diakonoff, 1976 Pammene piceae Komai, 1999 Pammene populana (Fabricius, 1787) Pammene pulchella Amsel, 1935 Pammene pullana Kuznetzov, 1986 Pammene purpureana (Constant, 1888) Pammene querceti (Gozmány, 1957) Pammene regiana (Zeller, 1849) Pammene rhediella (Clerck, 1759) Pammene salvana (Staudinger, 1859) Pammene shicotanica Kuznetzov in Danilevsky & Kuznetsov, 1968 Pammene soyoensis Bae & Park, 1998 Pammene spiniana (Duponchel in Godart, 1842) Pammene splendidulana (Guenée, 1845) Pammene subsalvana Kuznetzov, 1960 Pammene suspectana (Lienig & Zeller, 1846) Pammene tauriana Kuznetzov, 1960 Pammene trauniana (Denis & Schiffermüller, 1775) Pammene tricuneana (Kennel, 1900) Pammene tsugae Issiki in Issiki & Mutuura, 1961

Synonyms

Obsolete scientific names (junior synonyms and others) of Pammene are:^[3]

Encelis (lapsus) Eucelis Hübner, [1825] Encells (lapsus) Halonota Stephens, 1852 Hemene (lapsus) Hemerosia Stephens, 1852 Hemimene Hübner, 1825 Heusimene Stephens, 1834 Metasphaeroeca Fernald, 1908

Orchemia Guenée, 1845 Palla Billberg, 1820 (non Hübner, 1819: preoccupied) Pamene (lapsus) Pammena (lapsus) Phthoroblastis Lederer, 1859 Psudotomia Stephens, 1829 Pyrodes Guenée, 1845 (non Audinet-Serville, 1832: preoccupied) Sphaeroeca Meyrick, 1895 (non Lauterborn, 1894: preoccupied) Trycheris Guenée, 1845

The synonymy of Pammene has been subject to some confusion with its close relatives Cydia and Epinotia: Eucelis, Orchemia and Trycheris are sometimes^[4] listed as junior synonyms of the former, and Halonota has been listed^[5] as synonym of the latter. But the type species of Eucelis and Trycheris is Tortrix mediana (a junior synonym of P. aurana), that of Orchemia is O. gallicana (a junior synonym of P. gallicana), and that of Halonota is Pyralis populana (a junior synonym of P. populana). Thus, Eucelis, Halonota, Orchemia and Trycheris are all junior subjective synonym of the present genus.^[1]

Footnotes

1. Baixeras et al. (2009a)
2. Baixeras et al. (2009b)
3. Baixeras et al. (2009a), and see references in Savela (2005)
4. See e.g. references in Savela (2005)
5. See e.g. references in Savela (2009)

References

• Baixeras, J.; Brown, J.W. & Gilligan, T.M. (20091): Online World Catalogue of the Tortricidae – Pammene genus account. Version 1.3.1. Retrieved 2009-APR-20.
• Baixeras, J.; Brown, J.W. & Gilligan, T.M. (2009b): Online World Catalogue of the Tortricidae – Pammene species list. Version 1.3.1. Retrieved 2009-JAN-20.
• Savela, Markku (2005): Markku Savela's Lepidoptera and some other life forms – Pammene. Version of 2005-SEP-13. Retrieved 2010-APR-20.
• Savela, Markku (2009): Markku Savela's Lepidoptera and some other life forms – Epinotia. Version of 2009-JUL-07. Retrieved 2010-APR-19.