Parnassius imperator
Parnassius imperator | |
---|---|
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Lepidoptera |
Family: | Papilionidae |
Genus: | Parnassius |
Species: | P. imperator |
Binomial name | |
Parnassius imperator Oberthür, 1883 | |
Parnassius imperator is a high altitude butterfly which is found in Western China and Northeast India. It is a member of the generally white, high-elevation genus Parnassius, known as snow Apollos,[1] of the family Papilionidae, known as Swallowtails (although they lack tails).
It has over 40 subspecies, including the critically endangered Himalayan species P. i. augustus, commonly known as the Imperial Apollo.[2][3] Apollo Augustus, the ”Imperial” Apollo, appeared on Roman coins of the second century CE.[4]
When P. imperator come to rest, their wings spread horizontally so that their bodies lie close to the ground, lessening attacks by strong winds.[5]
Description
Subspecies P. i. augustus Fruhstorfer, 1903, from the 1907 book The Fauna of British India, including Ceylon and Burma – Butterflies, Volume II:[6]
- “♂ ♀. Upperside: creamy-white. Fore wing: costa and base closely irrorated with black scales; the usual short transverse medial and apical black bars across the cell, followed by irregularly sinuous, broad, transverse, dusky-black discal and postdiscal, complete bands that extend from the costa to the dorsum; the discal band very broad, below the apex of cell bent inwards and almost completely filling the base of interspaces 2 and 3; the terminal margin broadly dusky hyaline black. Hind wing: the dorsal margin up to the subcostal vein and downwards to a little above the tornus densely irrorated with black scales and with scattered long white hairs; an upper basal black-encircled crimson spot; another in the middle of interspace 7 and a third in interspace 5, the latter two centred with white ; beyond there is an irregular, sinuous, postdiscal dusky-black transverse band from costa to vein 4, posterior to which in interspaces 2 and 3 are two prominent round black subtornal spots, broadly centred with blue; lastly, the terminal margin narrowly dusky black. Cilia of both fore and hind wings prominently yellowish white. Underside similar; glassy markings similar, but mostly seen by transparency from above: faint indications of an additional crimson spot at base of cell and base of interspace 1. Antenna brownish black, obscurely annulated with brown ; head, thorax and abdomen black, the latter two studded with long white hairs.
- Exp. ♂ ♀ 79–84 mm. (3.1–3.35").
- Hab. Eastern Himalayas, on the boundary between Sikkim and Tibet, at altitudes between 15,000 and 16,000 feet.
- This smaller and darker form of P. imperator, Oberthür, from Eastern Tibet, just enters into our limits.”[6]
Range
“North-eastern India (Sikkim), western China (Xizang Zizhiqu (Tibet), Qinghai, Gansu, Sichuan (Szechwan) and Yunnan).”[7]
Status
P. imperator is “not known to be threatened as a species, but P. i. augustus is protected by law in India. It seems to be a common subalpine species in China.”[7] P. i. augustus was also included as a Schedule I subspecies in the Wildlife Protection Act, 1972.[2] P. i. augustus was listed as "Critically Endangered" in the 1994 The Red Data Book on Indian Animals: Butterflies of India.[3] P. imperator has been said to command a high price at European butterfly auctions in the past.[8]
Subspecies
There were twenty subspecies in 1985,[7] and more than twice that now included in the Global Names Index:[9][10]
- P. i. acalanatha Shinkai & Morita, 1995
- P. i. aino Bryk, 1932
- P. i. augustus Fruhstorfer, 1903
- P. i. aungsani Nose & Mikami, 1998
- P. i. cedermarki Bryk, 1934
- P. i. dominus Bang-Haas, 1934
- P. i. erlaensis Sugiyama, 1992
- P. i. evansi Bryk, 1932
- P. i. gigas Kotzsch, 1932
- P. i. haveli Kocman, 1995
- P. i. hoshinoi Kocman, 1999
- P. i. imperator Oberthür, 1883
- P. i. imperatrix Alph. 1897
- P. i. imposantus Schulte, 1991
- P. i. indra Korb 1997: Disputed validity.[11]
- P. i. interjungens Bryk 1932
- P. i. intermedius Rothschild 1909
- P. i. irmae Bryk, 1932
- P. i. jiyetiani Pierrat, 1990
- P. i. kameii Furumi & Sinkai, 1992
- P. i. karmapa Weiss & Michel, 1989
- P. i. kawasakii Sorimachi, 1992
- P. i. luxuriosus Mrácek & Schulte, 1990
- P. i. mahamayuri Shinkai & Morita, 1995
- P. i. milarepa Hamada, 2003
- P. i. musetta Bryk & Eisner, 1932
- P. i. musageta Grum-Grshimailo, 1891
- P. i. namchawarwanus Watanabe, 1995
- P. i. quaidami Kocman, 1995
- P. i. regina Bryk & Eisner, 1932
- P. i. regulus Bryk & Eisner, 1932
- P. i. rex Bang-Haas, 1928
- P. i. soi Sorimachi, 1997
- P. i. sultan Bryk & Eisner, 1932
- P. i. supremus Fruhstorfer, 1903
- P. i. takashii Ohya, 1990
- P. i. tara Shinkai, 1997
- P. i. titus Kocman, 1997
- P. i. tyrannus Bang-Haas, 1935
- P. i. uxoria Bang-Haas, 1935
- P. i. uxorius Bang-Haas, 1935
- P. i. vajramusti Oikawa, 1995
- P. i. venustus Stichel, 1906
Cited references
- ↑ Verghese, B. G. (1962). Himalayan endeavour. Times of India. p. 140.
- 1 2 The Wild Life (Protection) Act, 1972. Georg Thieme Verlag. p. 72.
- 1 2 Gupta, I. J.; Mondal, D. K.; Zoological Survey of India (1994). The Red Data Book on Indian Animals: Butterflies of India. Zoological Survey of India. p. 54. ISBN 978-81-8171-070-3.
- ↑ Graf, Fritz (16 October 2008). Apollo. Routledge. p. 102. ISBN 978-0-203-58171-1.
- ↑ Flora, fauna, and ecology of the Qinghai-Xizang (Tibet) Plateau. Science Press, Gordon & Breach Publishing Group. 1 January 1981. p. 1092. ISBN 978-0-677-60220-2.
- 1 2 Bingham, Charles Thomas (1907). The Fauna of British India, including Ceylon and Burma 2. London: Taylor & Francis. pp. 127–128.
- 1 2 3 Collins, N. Mark; Morris, Michael G. (1 January 1985). Threatened Swallowtail Butterflies of the World: The IUCN Red Data Book. International Union for Conservation of Nature. p. 37. ISBN 978-2-88032-603-6.
- ↑ Parenti, Umberto (1972). Butterflies and moths: a kaleidoscope of living colour; [adapted from the Italian of Umberto Parenti]. Orbis Publishing Ltd. p. 29.
- ↑ "Global Names Index". Global Names Index Beta. Global Biodiversity Information Facility and Enclopedia of Life. Retrieved 1 January 2013.
- ↑ "Taxonomy browser (Parnassius imperator)". NCBI Taxonomy Database. NCBI. Retrieved 1 January 2013.
- ↑ Häuser, Christoph L; Nekrutenko, Yuri P; (1998). "Comments on "Nomina Lepidopterorum nova" by S. K. Korb (Papilionidae, Nymphalidae)". Nota Lepidopterologica 21 (1): 74–84. ISSN 0342-7536.
Further reading
Parnassius imperator Sv Parnassius imperator provides further references and synonymy