Provanna

Provanna
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
(unranked):	clade Caenogastropoda
Superfamily:	Abyssochrysoidea
Family:	Provannidae
Genus:	Provanna Dall, 1918
Type species
Trichotropis (Provanna) lomana Dall, 1918

Provanna is a genus of sea snails, marine gastropod mollusks in the family Provannidae.^[1]

The species was named after Dall's first wife.

This is a very old genus with species already existing in Cretaceous seep deposits off Japan (93 - 100 million years ago). The Caribbean and the Pacific fauna were isolated about 3 million years ago by the closure of the Central American Isthmus.

Description

The apical whorls of the protoconch are lacking through decollation. The opening is then sealed with a calcareous plug. The sculpture of the ovate fusiform shell shows scattered sigmoidal axial ribs that are crossed by spiral cords. The whorls are broad.

Habitat

They are common in hydrothermal vents, hydrocarbon cold seeps and (more rarely) on deep-water food-falls, depending on the species. Most species feed on filamentous bacteria, while others feed on detritus.

Species

Species within the genus Provanna include:

• Provanna abyssalis Okutani & Fujikura, 2002^[2]
• Provanna admetoides Warén & Ponder, 1991^[3]
• † Provanna alexi Amano & Little, 2014
• † Provanna antiqua Squires, 1995
• Provanna buccinoides Warén & Bouchet, 1993^[4]
• Provanna chevalieri Warén & Bouchet, 2009^[5]
• Provanna clathrata Sasaki, Ogura, Watanabe & Fujikura, 2016
• Provanna glabra Okutani, Tsuchida & Fujikura, 1992^[6]
• Provanna goniata Warén & Bouchet, 1986^[7]
• † Provanna hirokoae Amano & Little, 2014
• Provanna ios Warén & Bouchet, 1986^[8]
• Provanna kuroshimensis Sasaki, Ogura, Watanabe & Fujikura, 2016
• Provanna laevis Warén & Ponder, 1991^[9]
• Provanna lomana (Dall, 1918)^[10]
• Provanna lucida Sasaki, Ogura, Watanabe & Fujikura, 2016
• Provanna macleani Warén & Bouchet, 1989^[11]
• † Provanna marshalli Saether, Little & K. A. Campbell, 2010
• Provanna muricata Warén & Bouchet, 1986^[12]
• † Provanna nakagawaensis Kaim, Jenkins & Hikida, 2009
• Provanna nassariaeformis Okutani, 1990^[13]
• Provanna pacifica (Dall, 1908)^[14]
• Provanna reticulata Warén & Bouchet, 2009^[15]
• Provanna sculpta Warén & Ponder, 1991^[16]
• Provanna segonzaci Warén & Ponder, 1991^[17]
• Provanna shinkaiae Okutani & Fujikura, 2002^[18]
• Provanna subglabra Sasaki, Ogura, Watanabe & Fujikura, 2016
• † Provanna tappuensis Kaim, Jenkins & Warén, 2008

†* Provanna urahoroensis Amano & Jenkins, 2013 

• Provanna variabilis Warén & Bouchet, 1986^[19]

Several fossil species also have been described, predominantly identified from hydrocarbon seep deposits (Saether et al., 2010), including:

• Provanna antiqua Squires & Goedert, 1995 ^[20]
• Provanna marshalli Saether, Little & Campbell, 2010
• Provanna nakagawaensis Kaim, Jenkins & Hikida, 2009
• Provanna tappuensis Kaim, Jenkins & Warén, 2008

There also are several fossil species known, but not formally described and named, from Central America, Japan, and possibly the Philippines (see Saether et al., 2010).

Species brought into synonymy

• Provanna marianensis Okutani & Fujikura, 1990: synonym of Desbruyeresia marianensis (Okutani & Fujikura, 1990)

References

1. Provanna Dall, 1918.  Retrieved through: World Register of Marine Species on 4 May 2010.
2. Provanna abyssalis Okutani & Fujikura, 2002.  Retrieved through: World Register of Marine Species on 4 May 2010.
3. Provanna admetoides Warén & Ponder, 1991.  Retrieved through: World Register of Marine Species on 4 May 2010.
4. Provanna buccinoides Warén & Bouchet, 1993.  Retrieved through: World Register of Marine Species on 4 May 2010.
5. Provanna chevalieri Warén & Bouchet, 2009.  Retrieved through: World Register of Marine Species on 4 May 2010.
6. Provanna glabra Okutani, Tsuchida & Fujikura, 1992.  Retrieved through: World Register of Marine Species on 4 May 2010.
7. Provanna goniata Warén & Bouchet, 1986.  Retrieved through: World Register of Marine Species on 4 May 2010.
8. Provanna ios Warén & Bouchet, 1986.  Retrieved through: World Register of Marine Species on 4 May 2010.
9. Provanna laevis Warén & Ponder, 1991.  Retrieved through: World Register of Marine Species on 4 May 2010.
10. Provanna lomana (Dall, 1918).  Retrieved through: World Register of Marine Species on 4 May 2010.
11. Provanna macleani Warén & Bouchet, 1989.  Retrieved through: World Register of Marine Species on 4 May 2010.
12. Provanna muricata Warén & Bouchet, 1986.  Retrieved through: World Register of Marine Species on 4 May 2010.
13. Provanna nassariaeformis Okutani, 1990.  Retrieved through: World Register of Marine Species on 4 May 2010.
14. Provanna pacifica (Dall, 1908).  Retrieved through: World Register of Marine Species on 4 May 2010.
15. Provanna reticulata Warén & Bouchet, 2009.  Retrieved through: World Register of Marine Species on 4 May 2010.
16. Provanna sculpta Warén & Ponder, 1991.  Retrieved through: World Register of Marine Species on 4 May 2010.
17. Provanna segonzaci Warén & Ponder, 1991.  Retrieved through: World Register of Marine Species on 4 May 2010.
18. Provanna shinkaiae Okutani & Fujikura, 2002.  Retrieved through: World Register of Marine Species on 4 May 2010.
19. Provanna variabilis Warén & Bouchet, 1986.  Retrieved through: World Register of Marine Species on 4 May 2010.
20. Steffen Kiel, New Records and Species of Molluscs from Tertiary Cold-Seep Carbonates in Washington State, USA; Journal of Paleontology, Vol. 80, No. 1 (Jan., 2006), pp. 121-137

• Healy, J. M. 1990. Taxonomic affinities of the deep-sea genus Provanna: new evidence from sperm structure; J. Molluscan Stud. 56: 119-122
• Spencer, H.G., Marshall, B.A. & Willan, R.C. 2009 Recent Mollusca. pp 196–219 in: Gordon, D.P. (Ed.), The New Zealand inventory of biodiversity. 1. Kingdom Animalia: Radiata, Lophotrochozoa, Deuterostomia. Canterbury University Press, Christchurch
• Saether, K.P., Little, C.T.S. & Campbell, K.A. (2010) A new fossil provannid gastropod from Miocene hydrocarbon seep deposits, East Coast Basin, North Island, New Zealand. Acta Palaeontologica Polonica 55, 507–517.

External links