Monofenestrata

Monofenestratans
Temporal range:
Middle JurassicLate Cretaceous, 166–66 Ma
Fossil specimen of Darwinopterus modularis
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Order: Pterosauria
Clade: Pterodactylomorpha
Clade: Monofenestrata
et al., 2010
Subgroups

The Monofenestrata are an unranked group of pterosaurs that includes the family Wukongopteridae and the suborder Pterodactyloidea.[1]

The clade Monofenestrata was in 2009/2010 defined as the group consisting of Pterodactylus and all species sharing with Pterodactylus the synapomorphy, shared derived trait, of an external nostril confluent with the antorbital fenestra, the major skull opening on the side of the snout. The name is derived from Greek monos, "single", and Latin fenestra, "window". The concept was inspired by the discovery of Darwinopterus, a species combining a pterodactyloid-type skull with a more basal build of the remainder of the body.[1] The Darwinoptera, a primitive subgroup of monofenestratans showing this transitional anatomy, was also named for Darwinopterus and defined as all descendants of its common ancestor with Pterorhynchus.[2]

The earliest known monofenestrate fossils have been found in the Stonesfield Slate formation of the United Kingdom, which dates to the Bathonian stage of the Middle Jurassic, dated to about 166 million years ago.[3]

Below is a cladogram showing the results of a phylogenetic analysis presented by Andres, Clark & Xu, 2014. This study found the two traditional groupings of ctenochasmatoids and kin as an early branching group, with all other pterodactyloids grouped into the Eupterodactyloidea.[2]

 Pterodactylomorpha 

Sordes


 Monofenestrata 
Darwinoptera

Pterorhynchus



Wukongopteridae



Pterodactyliformes 

Changchengopterus


 Caelidracones 

Anurognathidae



Pterodactyloidea






References

  1. 1 2 Lü, J., Unwin, D.M., Jin, X., Liu, Y. and Ji, Q. (2010). "Evidence for modular evolution in a long-tailed pterosaur with a pterodactyloid skull." Proceedings of the Royal Society B, 277(1680): 383-389. doi:10.1098/rspb.2009.1603 PMID 19828548
  2. 1 2 Andres, B.; Clark, J.; Xu, X. (2014). "The Earliest Pterodactyloid and the Origin of the Group". Current Biology. doi:10.1016/j.cub.2014.03.030.
  3. Buffetaut, E. and Jeffrey, P. (2012). "A ctenochasmatid pterosaur from the Stonesfield Slate (Bathonian, Middle Jurassic) of Oxfordshire, England." Geological Magazine, (advance online publication) doi:10.1017/S0016756811001154
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