Staphylinoidea

Staphylinoidea

Devil's coach horse beetle, Ocypus olens
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Suborder:	Polyphaga
Infraorder:	Staphyliniformia
Superfamily:	Staphylinoidea Latreille, 1802
Families
See text.

Wikispecies has information related to: Staphylinoidea

Staphylinoidea is a superfamily of beetles. It is a very large and diverse group with worldwide distribution.

Characteristics

Most staphylinoids are middle-sized or small beetles with a pair of shortened elytra. The hind wings have no accessory posterior ridge (locking device), no wedge cell and apical cell. Abdominal 8th segment not entirely invaginated in the 7th. Head mostly without coronal suture.^[1]

Systematics and evolution

Staphylinoidea contains the following subgroups:^[2]^[3]

Agyrtidae C.G. Thomson 1859
Hydraenidae Mulsant 1844
Leiodidae Fleming 1821 = (Anisotomidae)
Ptiliidae Erichson 1845
- Cephaloplectinae or Limulodidae Sharp 1883
Scydmaenidae Leach 1815
Silphidae Latreille 1807 (carrion beetles)
Staphylinidae Latreille 1802 (rove beetles)

The unambiguous fossil record dates back to Triassic, and an early Mesozoic origin of the group is probable.^[4]

References

↑ M.Hansen. Phylogeny and classification of the staphyliniform beetle families (Coleoptera). Biologiske Skrifter 48, Copenhagen, 1997
↑ Beutel, R. G. and Leschen, R.A.B. 2005. Phylogenetic analysis of Staphyliniformia (Coleoptera) based on characters of larvae and adults. Systematic Entomology
↑ J. F. Lawrence and A. F. Newton, Jr. 1995. Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names). pp. 779-1006 In: J. Pakaluk & S. A. Slipinski (Eds.): Biology, Phylogeny, and Classification of Coleoptera: Papers Celebrating the 80th Birthday of Roy A. Crowson. Museum i Instytut Zoologii PAN, Warszawa
↑ Grimaldi, D. and Engel, M.S. (2005). Evolution of the Insects. Cambridge University Press

Extant Coleoptera families

Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Subclass: Pterygota
Infraclass: Neoptera
Superorder: Endopterygota

Suborder Archostemata

Crowsoniellidae (Crowsoniella relicta) Cupedidae (reticulated beetles) Jurodidae (Sikhotealinia zhiltzovae) Micromalthidae (telephone-pole beetle) Ommatidae

Suborder Adephaga

Extant families	Amphizoidae (trout-stream beetles) Aspidytidae Carabidae (ground beetles) Dytiscidae (predaceous diving beetles) Gyrinidae (whirligig beetles) Haliplidae (crawling water beetles) Hygrobiidae Meruidae (Meru phyllisae) Noteridae (burrowing water beetles) Rhysodidae (wrinkled bark beetles) Trachypachidae (false ground beetles)

Extinct families	Colymbothetidae Coptoclavidae Liadytidae Parahygrobiidae Triaplidae

Suborder Myxophaga

Hydroscaphidae (skiff beetles) Lepiceridae Sphaeriusidae Torridincolidae

Suborder Polyphaga

Bostrichiformia

Bostrichoidea	Anobiidae (furniture beetles, death watch beetles, spider beetles) Bostrichidae (auger beetles) Dermestidae (skin beetles) Jacobsoniidae (Jacobson's beetles) Nosodendridae (wounded-tree beetles)

Derodontoidea	Derodontidae (tooth-necked fungus beetles)

Cucujiformia

Chrysomeloidea	Cerambycidae (longhorn beetles) Chrysomelidae (leaf beetles) Disteniidae Megalopodidae Orsodacnidae Oxypeltidae Vesperidae

Cleroidea	Acanthocnemidae (Acanthocnemus nigricans) Chaetosomatidae Cleridae (checkered beetles) Melyridae (soft-wing flower beetles) Phloiophilidae (Phloiophilus edwardsi) Phycosecidae Prionoceridae Trogossitidae (bark-gnawing beetles)

Cucujoidea	Alexiidae Biphyllidae (false skin beetles) Boganiidae Bothrideridae (dry bark beetles) Byturidae (fruitworm beetles) Cavognathidae Cerylonidae (minute bark beetles) Coccinellidae (lady beetles, or God's cows) Corylophidae (minute fungus beetles) Cryptophagidae (silken fungus beetles) Cucujidae (flat bark beetles) Cyclaxyridae Discolomatidae Endomychidae (handsome fungus beetles) Erotylidae (pleasing fungus beetles) Helotidae Hobartiidae Kateretidae (short-winged flower beetles) Laemophloeidae (lined flat bark beetles) Lamingtoniidae (Lamingtonium binnaberrense) Latridiidae (minute brown scavenger beetles) Monotomidae (root-eating beetles) Myraboliidae Nitidulidae (sap beetles) Passandridae (parasitic flat bark beetles) Phalacridae (shining flower beetles) Phloeostichidae Propalticidae Protocucujidae Silvanidae (silvanid flat bark beetles) Smicripidae (palmetto beetles) Sphindidae (dry-fungus beetles)

Curculionoidea (weevils)	Anthribidae (fungus weevils) Attelabidae (leaf-rolling weevils) Belidae (primitive weevils) Brentidae (straight snout weevils, New York weevil) Caridae Curculionidae (true weevils, bark beetles, ambrosia beetles) Nemonychidae (pine flower weevils)

Lymexyloidea	Lymexylidae (ship-timber beetles)

Tenebrionoidea	Aderidae (ant-like leaf beetles) Anthicidae (ant-like flower beetles) Archeocrypticidae (cryptic fungus beetles) Boridae (conifer bark beetles) Chalcodryidae Ciidae (minute tree-fungus beetles) Melandryidae (false darkling beetles) Meloidae (blister beetles) Mordellidae (tumbling flower beetles) Mycetophagidae (hairy fungus beetles) Mycteridae (palm and flower beetles) Oedemeridae (false blister beetle) Perimylopidae, or Promecheilidae Prostomidae (jugular-horned beetles) Pterogeniidae Pyrochroidae (fire-coloured beetles) Pythidae (dead log bark beetles) Ripiphoridae (wedge-shaped beetles) Salpingidae (narrow-waisted bark beetles) Scraptiidae (false flower beetles) Stenotrachelidae (false longhorn beetles) Synchroidae (synchroa bark beetles) Tenebrionidae (darkling beetles) Tetratomidae (polypore fungus beetles) Trachelostenidae Trictenotomidae Ulodidae Zopheridae (ironclad beetles, cylindrical bark beetles)

Elateriformia

Buprestoidea	Buprestidae (jewel beetles, or metallic wood-boring beetles) Schizopodidae

Byrrhoidea	Byrrhidae (pill beetles) Callirhipidae (cedar beetles) Chelonariidae (turtle beetles) Cneoglossidae Dryopidae (long-toed water beetles) Elmidae (riffle beetles) Eulichadidae (forest stream beetles) Heteroceridae (variegated mud-loving beetles) Limnichidae (minute mud beetles) Lutrochidae (travertine beetles) Psephenidae (water-penny beetles) Ptilodactylidae

Dascilloidea	Dascillidae (soft bodied plant beetles) Rhipiceridae (cicada beetle, cicada parasite beetles)

Elateroidea	Anischiidae Artematopodidae (soft-bodied plant beetles) Brachypsectridae (Texas beetles) Cantharidae (soldier beetles) Cerophytidae (rare click beetles) Drilidae Elateridae (click beetles) Eucnemidae (false click beetles) Lampyridae (fireflies) Lycidae (net-winged beetles) Omalisidae Omethidae (false fireflies) Phengodidae (glowworm beetles, long-lipped beetles) Plastoceridae (Plastocerus angulosus) Podabrocephalidae Rhagophthalmidae Rhinorhipidae (Rhinorhipus tamborinensis) Throscidae (false metallic wood-boring beetles)

Scirtoidea	Clambidae Decliniidae (Declinia relicta) Eucinetidae (plate-thigh beetles) Scirtidae

Scarabaeiformia

Scarabaeoidea

Staphyliniformia

Histeroidea	Histeridae (clown beetles) Sphaeritidae (false clown beetles) Synteliidae

Hydrophiloidea	Epimetopidae Georissidae (minute mud-loving beetles) Helophoridae Hydrochidae Hydrophilidae (water scavenger beetles) Spercheidae

Staphylinoidea	Agyrtidae (primitive carrion beetles) Hydraenidae Leiodidae (round fungus beetles) Ptiliidae (feather-winged beetles) Scydmaenidae (ant-like stone beetles) Silphidae (carrion beetles) Staphylinidae (rove beetles)

† Suborder Protocoleoptera

Ademosyndidae Labradorocoleoidae Oborocoleoidae Permocupedidae Permosynidae Taldycupedidae Tshekardocoleoidae

List of subgroups of the order Coleoptera

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