Triplophysa
Triplophysa | |
---|---|
Triplophysa stoliczkae | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Actinopterygii |
Order: | Cypriniformes |
Family: | Nemacheilidae |
Genus: | Triplophysa Rendahl (de), 1933 |
Type species | |
Nemacheilus hutjertjuensis Rendahl, 1933 |
Triplophysa is a large genus of stone loaches,[1][2][3] most of which are found in and around the Qinghai-Tibet Plateau (China).[4]
Currently Triplophysa is a mixed assemblage of species. Some lineages have been identified and treated as subgenera (Hedinichthys, Qinghaichthys, Labiatophysa, Indotriplophysa, and Tarimichthys),[1] but following Kottelat (2012)[1] and the Catalog of Fishes[3] are treated as genera in Wikipedia. FishBase, however, includes these in Triplophysa.[2]
Ecology
Triplophysa zhaoi holds the record for the lowest altitude for Asian fish: it is found at 50 m (160 ft) below sea level in swamps of the Lükqün oasis, in the Turpan depression, Xinjiang. In the other end, Triplophysa stoliczkae holds the record altitude for Asian fish: it is found at 5,200 m (17,100 ft) above sea level in hot springs near the Longmu Lake in western Tibet.[1] Triplophysa dalaica has been used as model species to study adaptation to high-altitude hypoxia, and 13 positively selected genes involved in hypoxia response have been identified.[5] Some species are blind troglobites.[4][6]
Species
There are currently 125 recognized species in this genus:[1]
- Triplophysa alexandrae Prokofiev, 2001
- Triplophysa aliensis (Y. F. Wu & S. Q. Zhu, 1979)
- Triplophysa altipinnis Prokofiev, 2003
- Triplophysa aluensis W. X. Li & Z. G. Zhu, 2000
- Triplophysa angeli (P. W. Fang, 1941)
- Triplophysa anterodorsalis S. Q. Zhu & W. X. Cao, 1989
- Triplophysa aquaecaeruleae Prokofiev, 2001
- Triplophysa arnoldii Prokofiev, 2006
- Triplophysa bashanensis T. Q. Xu & Ka. Fe. Wang, 2009
- Triplophysa bellibarus (T. L. Tchang, T. H. Yueh & H. C. Hwang, 1963)
- Triplophysa bleekeri (Sauvage & Dabry de Thiersant, 1874)
- Triplophysa brachyptera (Herzenstein, 1888)
- Triplophysa brahui (Zugmayer, 1912)
- Triplophysa brevibarba R. H. Ding, 1993
- Triplophysa brevicauda (Herzenstein, 1888)
- Triplophysa cakaensis W. X. Cao & S. Q. Zhu, 1988
- Triplophysa chondrostoma (Herzenstein, 1888)
- Triplophysa coniptera (Turdakov, 1954)
- Triplophysa crassilabris R. H. Ding, 1994
- Triplophysa cuneicephala (T. H. Shaw & T. L. Tchang, 1931)
- Triplophysa dalaica (Kessler, 1876)
- Triplophysa daqiaoensis R. H. Ding, 1993
- Triplophysa dongganensis Jian Yang, 2013 [7]
- Triplophysa dorsalis (Kessler, 1872) (Gray loach)
- Triplophysa dorsonotata (Kessler, 1879)
- Triplophysa drassensis (Tilak, 1990)
- Triplophysa farwelli (Hora, 1935)
- Triplophysa fengshanensis J. H. Lan, 2013 [7]
- Triplophysa flavicorpus J. X. Yang, X. Y. Chen & J. H. Lan, 2004
- Triplophysa furva S. Q. Zhu, 1992
- Triplophysa fuxianensis J. X. Yang & X. L. Chu, 1990
- Triplophysa gejiuensis (X. L. Chu & Y. R. Chen, 1979)
- Triplophysa gerzeensis W. X. Cao & S. Q. Zhu, 1988
- Triplophysa gracilis (F. Day, 1877)
- Triplophysa grahami (Regan, 1906)
- Triplophysa griffithi (Günther, 1868)
- Triplophysa gundriseri Prokofiev, 2002
- Triplophysa hexiensis (T. Q. Zhao & X. T. Wang, 1988)
- Triplophysa heyangensis S. Q. Zhu, 1992
- Triplophysa hialmari Prokofiev, 2001
- Triplophysa hsutschouensis (Rendahl (de), 1933)
- Triplophysa huanglongensis Gao, 1992
- Triplophysa huanjiangensis Jian Yang, T. J. Wu & J. H. Lan, 2011
- Triplophysa huapingensis L. P. Zheng, J. X. Yang & X. Y. Chen, 2012 [8]
- Triplophysa hutjertjuensis (Rendahl (de), 1933)
- Triplophysa intermedia (Kessler, 1876)
- Triplophysa jianchuanensis L. P. Zheng, L. N. Du, X. Y. Chen & J. X. Yang, 2010 (Jianchuan stone loach)
- Triplophysa jiarongensis Y. Lin, Chao Li & J. K. Song, 2012 [9]
- Triplophysa kafirnigani (Turdakov, 1948)
- Triplophysa kullmanni Bănărescu & Nalbant, 1975
- Triplophysa kungessana (Kessler, 1879)
- Triplophysa lacusnigri (L. S. Berg, 1928)
- Triplophysa lacustris J. X. Yang & X. L. Chu, 1990
- Triplophysa langpingensis Jian Yang, 2013 [7]
- Triplophysa laterimaculata J. L. Li, N. F. Liu & J. X. Yang, 2007
- Triplophysa laticeps W. Zhou & G. H. Cui, 1997
- Triplophysa leptosoma (Herzenstein, 1888) [10]
- Triplophysa lihuensis T. J. Wu, Jian Yang & J. H. Lan, 2012 [11]
- Triplophysa lingyunensis (J. W. Liao, D. Z. Wang & Z. F. Luo, 1997)
- Triplophysa lixianensis C. L. He, Z. B. Song & E. Zhang, 2008
- Triplophysa longianguis Y. F. Wu & C. Z. Wu, 1984
- Triplophysa longibarbata (Y. R. Chen, J. X. Yang, Sket & Aljančič, 1998)
- Triplophysa longipectoralis L. P. Zheng, L. N. Du, X. Y. Chen & J. X. Yang, 2009
- Triplophysa longliensis Q. Ren, J. X. Yang & X. Y. Chen, 2012 [12]
- Triplophysa macrocephala Jian Yang, T. J. Wu & J. X. Yang, 2012 [13]
- Triplophysa macromaculata J. X. Yang, 1990
- Triplophysa macrophthalma S. Q. Zhu & Q. Z. Guo, 1985
- Triplophysa markehenensis (S. Q. Zhu & Y. F. Wu, 1981)
- Triplophysa marmorata (Heckel, 1838)
- Triplophysa microphthalma (J. W. Liao & D. Z. Wang, 1997)
- Triplophysa microphysa (P. W. Fang, 1935)
- Triplophysa microps (Steindachner, 1866)
- Triplophysa moquensis R. H. Ding, 1994
- Triplophysa nandanensis J. H. Lan, J. X. Yang & Y. R. Chen, 1995
- Triplophysa nanpanjiangensis (S. Q. Zhu & W. X. Cao, 1988)
- Triplophysa nasobarbatula D. Z. Wang & D. J. Li, 2001
- Triplophysa ninglangensis Y. F. Wu & C. Z. Wu, 1988
- Triplophysa nujiangensa X. Y. Chen, G. H. Cui & J. X. Yang, 2004
- Triplophysa obscura X. T. Wang, 1987
- Triplophysa obtusirostra Y. F. Wu & C. Z. Wu, 1988
- Triplophysa orientalis (Herzenstein, 1888)
- Triplophysa papillosolabiata (Kessler, 1879)
- Triplophysa pappenheimi (P. W. Fang, 1935)
- Triplophysa paradoxa (Turdakov, 1955)
- Triplophysa parvus Z. M. Chen, W. X. Li & J. X. Yang, 2009
- Triplophysa pedaschenkoi (L. S. Berg, 1931)
- Triplophysa polyfasciata R. H. Ding, 1996
- Triplophysa pseudoscleroptera (S. Q. Zhu & Y. F. Wu, 1981)
- Triplophysa pseudostenura C. L. He, E. Zhang & Z. B. Song, 2012 [14]
- Triplophysa qilianensis W. J. Li, X. C. Chen & Y. P. Hu, 2015 [15]
- Triplophysa qiubeiensis W. X. Li & H. F. Yang, 2008
- Triplophysa robusta (Kessler, 1876)
- Triplophysa rosa X. Y. Chen & J. X. Yang, 2005 [4]
- Triplophysa scapanognatha Prokofiev, 2007
- Triplophysa scleroptera (Herzenstein, 1888)
- Triplophysa sellaefer (Nichols, 1925)
- Triplophysa sewerzowi (A. M. Nikolskii, 1938) (Severtsov’s loach)
- Triplophysa shaanxiensis J. X. Chen, 1987
- Triplophysa shehensis Menon, 1987
- Triplophysa shilinensis Y. R. Chen & J. X. Yang, 1992
- Triplophysa shiyangensis (T. Q. Zhao & X. T. Wang, 1983)
- Triplophysa siluroides (Herzenstein, 1888)
- Triplophysa stenura (Herzenstein, 1888)
- Triplophysa stewarti (Hora, 1922)
- Triplophysa stoliczkae (Steindachner, 1866) (Tibetan stone loach) [10]
- Triplophysa strauchii (Kessler, 1874) (Spotted thicklip loach)
- Triplophysa tanggulaensis (S. Q. Zhu, 1982)
- Triplophysa tianeensis X. Y. Chen, G. H. Cui & J. X. Yang, 2004
- Triplophysa tibetana (Regan, 1905)
- Triplophysa turpanensis Y. F. Wu & C. Z. Wu, 1992
- Triplophysa ulacholica (Anikin, 1905) (Issyk-kul naked loach)
- Triplophysa uranoscopus (Kessler, 1872)
- Triplophysa venusta S. Q. Zhu & W. X. Cao, 1988
- Triplophysa waisihani L. Cao & E. Zhang, 2008
- Triplophysa wuweiensis (S. C. Li & S. Y. Chang, 1974)
- Triplophysa xiangshuingensis W. X. Li, 2004
- Triplophysa xiangxiensis (G. Y. Yang, F. X. Yuan & Y. M. Liao, 1986)
- Triplophysa xichangensis S. Q. Zhu & W. X. Cao, 1989
- Triplophysa xingshanensis (G. R. Yang & C. X. Xie, 1983)
- Triplophysa xiqiensis R. H. Ding & Q. Lai, 1996
- Triplophysa yajiangensis S. L. Yan, Z. Y. Sun & Y. S. Guo, 2015 [10]
- Triplophysa yaopeizhii T. Q. Xu, C. G. Zhang & B. Cai, 1995
- Triplophysa yunnanensis J. X. Yang, 1990
- Triplophysa zhaoi Prokofiev, 2006
- Triplophysa zhenfengensis D. Z. Wang & D. J. Li, 2001
References
Wikimedia Commons has media related to Triplophysa. |
- 1 2 3 4 5 Kottelat M (2012). "Conspectus cobitidum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei)" (PDF). Raffles Bulletin of Zoology, Suppl 26: 1–199.
- 1 2 Froese, Rainer, and Daniel Pauly, eds. (2015). Species of Triplophysa in FishBase. October 2015 version.
- 1 2 Eschmeyer, W.N. (2 June 2015). "Catalog of Fishes". California Academy of Sciences.
- 1 2 3 Chen X.-Y., Yang J.-X. (2005). "Triplophysa rosa sp. nov.: a new blind loach from China". Journal of Fish Biology 66 (3): 599–608. doi:10.1111/j.0022-1112.2005.00622.x.
- ↑ Wang, Y., Yang, L., Wu, B., Song, Z. & He, S. (2015). "Transcriptome analysis of the plateau fish (Triplophysa dalaica): Implications for adaptation to hypoxia in fishes". Gene 565 (2): 211–220. doi:10.1016/j.gene.2015.04.023.
- ↑ Romero, A., Zhao, Y. & Chen, X. (2009). "The hypogean fishes of China". Environmental Biology of Fishes 86: 211–278. doi:10.1007/s10641-009-9441-3.
- 1 2 3 Lan, J., Gan. X., Wu. T. & Yang, J. (2013): Cave Fishes of Guangxi, China. Science Press, Beijing. 1-266.
- ↑ Zheng L.-P., Yang J.-X., Chen X.-Y. (2012). "A new species of Triplophysa (Nemacheilidae: Cypriniformes), from Guangxi, southern China". Journal of Fish Biology 80 (4): 831–841. doi:10.1111/j.1095-8649.2012.03227.x.
- ↑ Lin Y., Li C., Song J.-K. (2012). "A new species of troglobitic loach (Cypriniformes, Balitoridae), Triplophysa jiarongensis, from Guizhou, China". Acta Zootaxonomica Sinica 37 (3): 640–647.
- 1 2 3 Yan S.-L., Sun Z.-Y., Guo Y.-S. (2015). "A new species of Triplophysa Rendahl (Cypriniformes, Nemacheilidae) from Sichuan Province, China". Zoological Research 36 (5): 299–304.
- ↑ Wu T.-J., Yang J., Lan J.-H. (2012). "A new blind loach, Triplophysa lihuensis (Teleostei: Balitoridae), from Guangxi, China" (PDF). Zoological Studies 51 (6): 874–880.
- ↑ Ren Q., Yang J.-X., Chen X.-Y. (2012). "A new species of the genus Triplophysa (Cypriniformes: Nemacheilidae), Triplophysa longliensis sp. nov, from Guizhou, China" (PDF). Zootaxa 3586: 187–194.
- ↑ Yang J., Wu T.-J., Yang J.-X. (2012). "A new cave-dwelling loach, Triplophysa macrocephala (Teleostei: Cypriniformes: Balitoridae), from Guangxi, China". Environmental Biology of Fish 93 (2): 169–175. doi:10.1007/s10641-011-9901-4.
- ↑ He C., Zhang E, Song Z. (2012). "Triplophysa pseudostenura, a new nemacheiline loach (Cypriniformes: Balitoridae) from the Yalong River of China" (PDF). Zootaxa 3586: 272–280.
- ↑ Li W.-J., Chen X.-C., Hu Y.-P. (2015). "A new species of the genus Triplophysa (Nemacheilinae), Triplophysa qilianensis sp. nov, from Qinghai, China". Zootaxa 3905 (3): 418–424. doi:10.11646/zootaxa.3905.3.7.