Anteophthalmosuchus

Anteophthalmosuchus
Temporal range: Early Cretaceous
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Superorder: Crocodylomorpha
Family: Goniopholididae
Genus: Anteophthalmosuchus
Salisbury & Naish, 2011
Type species
Anteophthalmosuchus hooleyi
Salisbury & Naish, 2011
Species
  • A. escuchae Buscalioni et al., 2013
  • A. hooleyi Salisbury & Naish, 2011

Anteophthalmosuchus (meaning "forward-pointing eye crocodile") is an extinct genus of goniopholidid mesoeucrocodylian from the Early Cretaceous of southern England and eastern Spain.[1][2][3]

It is known from a single holotype specimen from the Wealden Group of the Isle of Wight that includes a well-preserved skull and partial skeleton. This specimen has been known since 1904 and was identified as the "Tie Pits specimen" or the "Hooley specimen" after Reginald Walter Hooley, an amateur paleontologist who had described it in 1905. Hooley had originally attributed the specimen to the previously named species Goniopholis crassidens.[4]

In 2011, Hooley's specimen was redescribed as a distinct genus and species of goniopholidid called Anteophthalmosuchus hooleyi. The genus name means "forward-pointing eye crocodile" because the specimen's eye sockets are positioned high on the skull and angle forward rather than to the side as in most other flat-skulled crocodyliforms, and the species name honors Hooley. A second, smaller skull was identified as a possible juvenile specimen of Anteophthalmosuchus. Features that distinguish A. hooleyi from Goniopholis crassidens include the lack of a hole in the lower jaw called the mandibular fenestra, very wide supratemporal fenestrae (openings) on the skull table, and a bone above the eye socket called the palpebral that is small and does not extend over the socket as in some other goniopholidids.[4]

At an estimated 3.5 metres (11 ft) in length, A. hooleyi would have been the largest crocodyliform in the Wealden faunal assemblage, larger than the contemporaneous species Hylaeochampsa vectiana, Leiokarinosuchus brookensis, and Vectisuchus leptognathus.[4]

A second species, Anteophthalmosuchus escuchae, was first described and named by A.D. Buscalioni, L. Alcalá, E. Espílez and L. Mampel in 2013. It is known solely from the holotype AR-1-1097 which consists of a partial skull. It was collected from the early Albian-aged Escucha Formation, at Santa Maria Mine located in the municipality of Ariño, Teruel Province, of Aragon, along with the closely related Hulkepholis plotos.[3]

Phylogeny

Anteophthalmosuchus hooleyi was included in a phylogenetic analysis of goniopholidids that was published soon before the specimen was redescribed. It was found to be most closely related to a specimen called "Dollo's goniopholidid", and the two were found to be the closest relatives of a specimen called "Hulke's goniopholidid", now named Goniopholis willetti. Below is a cladogram from that analysis:[5]

Neosuchia
Atoposauridae

Theriosuchus pusillus



Theriosuchus guimarotae






Rugosuchus




Bernissartia



Eusuchia






Stolokrosuchus





Tethysuchia



Thalattosuchia



Goniopholididae

Calsoyasuchus valliceps





"Goniopholis" phuwiangensis




Eutretauranosuchus delfi



"Sunosuchus" junggarensis





Sunosuchus miaoi



Sunosuchus thailandicus






Siamosuchus phuphokensis




Amphicotylus lucasii




Denazinosuchus kirtlandicus



Nannosuchus gracilidens





Hulke’s goniopholidid




Hooley’s goniopholidid



Dollo’s goniopholidid




Goniopholis

Goniopholis baryglyphaeus




Goniopholis kiplingi



Goniopholis simus














References

  1. Steven W. Salisbury and Darren Naish (2011). "Crocodilians". In Batten, D. J. (eds). English Wealden Fossils. The Palaeontological Association (London). pp. 305–369.
  2. Naish, Darren (2 December 2011). "The Wealden Bible: English Wealden Fossils, 2011". Tetrapod Zoology. Scientific American Blogs. Retrieved 5 December 2011.
  3. 1 2 Buscalioni, A.D.; Alcalá, L.; Espílez, E. & Mampel, L. (2013). "European Goniopholididae from the Early Albian Escucha Formation in Ariño (Teruel, Aragón, España)". Spanish Journal of Palaeontology 28 (1): 103–122.
  4. 1 2 3 Naish, Darren (24 September 2012). "In pursuit of Early Cretaceous crocodyliforms in southern England: ode to Goniopholididae". Tetrapod Zoology. Scientific American Blogs. Retrieved 24 September 2012.
  5. De Andrade, M. B.; Edmonds, R.; Benton, M. J.; Schouten, R. (2011). "A new Berriasian species of Goniopholis (Mesoeucrocodylia, Neosuchia) from England, and a review of the genus". Zoological Journal of the Linnean Society 163: S66. doi:10.1111/j.1096-3642.2011.00709.x.
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