2013 in paleontology

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Paleontology, palaeontology or [palæontology (from Greek: paleo, "ancient"; ontos, "being"; and logos, "knowledge") is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.^[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because mankind has encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2013.

Plants

Angiosperms

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Atli[2]	Gen. et sp. nov.	Valid	Smith et al.	Late Cretaceous (Campanian)		Canada	A ranunculalean known from a stem with lianoid anatomy. The type species is Atli morinii.
Bambusiculmus[3]	Gen. et 2 sp. nov.	Valid	Wang & Zhou in Wang et al.	Miocene	Dajie Formation	China	A bamboo. The type species is Bambusiculmus angustus; genus also includes Bambusiculmus latus.
Bambusium angustifolia[3]	Sp. nov.	Valid	Wang & Zhou in Wang et al.	Miocene	Dajie Formation	China	A bamboo, a species of Bambusium.
Bambusium latifolia[3]	Sp. nov.	Valid	Wang & Zhou in Wang et al.	Miocene	Dajie Formation	China	A bamboo, a species of Bambusium.
Bombacacioxylon[4]	Gen. et sp. nov.	Valid	Mehrotra et al.	Late Miocene to early Pliocene	Tipam Group	India	A possible relative of Bombax. The type species is Bombacacioxylon tertiarum.
Firmiana oligocenica[5]	Sp. nov.	Valid	Srivastava & Mehrotra	Oligocene (Chattian)	Tikak Parbat Formation	India	A species of Firmiana.
Guadua morronei[6]	Sp. nov.	Valid	Brea, Zucol & Franco	Pliocene–Pleistocene	Ituzaingó Formation	Argentina	A bamboo, a species of Guadua.
Iterophyllum[7]	Gen. et sp. nov.	Valid	Barral et al.	Early Cretaceous (late Barremian)		Spain	A basal eudicot. The type species is Iterophyllum lobatum.
Jaguariba wiersemana[8]	Gen. nov. et sp. nov.	Valid	Coiffard, Mohr & Bernardes-de-Oliveira	Early Cretaceous	Crato Formation	Brazil	Fossil Nymphaeales.
Palmoxylon vastanensis[9]	Sp. nov.	Valid	Prasad et al.	Eocene	Cambay Shale Formation	India	A palm tree, a species of Palmoxylon.
Paranephelium makumensis[5]	Sp. nov.	Valid	Srivastava & Mehrotra	Oligocene (Chattian)	Tikak Parbat Formation	India	A species of Paranephelium.
Poeciloneuron preindicum[10]	Sp. nov.	Valid	Srivastava & Mehrotra	Late Oligocene		India	A species of Poeciloneuron.
Potomacapnos[11]	Gen. et sp. nov.	Valid	Jud & Hickey	Early Cretaceous (Aptian)	Potomac Group	United States	A close relative of ranunculalean eudicots. The type species is Potomacapnos apeleutheron.
Pterygota palaeoalata[5]	Sp. nov.	Valid	Srivastava & Mehrotra	Oligocene (Chattian)	Tikak Parbat Formation	India	A species of Pterygota.
Sapindus palaeoemarginatus[5]	Sp. nov.	Valid	Srivastava & Mehrotra	Oligocene (Chattian)	Tikak Parbat Formation	India	A species of Sapindus.
Spixiarum kipea[12]	Gen. nov. et sp. nov.	Valid	Coiffard, Mohr & Bernardes-de-Oliveira	Early Cretaceous	Crato Formation	Brazil	Fossil Aroid.
Virola dominicana[13]	Sp. nov.	Valid	Poinar & Steeves	Cenozoic (Eocene or Miocene)		Dominican Republic	A member of Myristicaceae found in Dominican amber, a species of Virola.

Other seed plants

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Donponoxylon[14]	Gen. et 2 sp. nov.	Valid	Tidwell, Britt & Wright	Middle to Late Jurassic		Australia  New Zealand	A seed plant of uncertain phylogenetic placement. The type species is Donponoxylon bennettii; genus also contains Donponoxylon jacksonii.
Friedsellowia gracilifolia[15]	Gen. nov. et sp. nov.	Valid	Löwe et al.	Early Cretaceous	Crato Formation	Brazil	A gnetophyte. The type species of a new genus.
Itajuba[16]	Gen. et sp. nov	Valid	Ricardi-Branco et al.	Early Cretaceous (Aptian or Albian)	Crato Member of the Santana Formation	Brazil	A member of Gnetales. The type species is Itajuba yansanae.
Pagiophyllum heshanense[17]	Sp. nov.	Valid	Liu & Yao	Permian		China	A species of Pagiophyllum.
Pararaucaria delfueyoi[18]	Sp. nov.	Valid	Escapa et al.	Late Jurassic	Cañadón Calcáreo Formation	Argentina	A member of Coniferales belonging to the family Cheirolepidiaceae, a species of Pararaucaria.
Schizolepidopsis canicularis[19]	Sp. nov.	Valid	Leslie et al.	Early Cretaceous (probably Aptian-Albian)	Tevshiin Govi Formation	Mongolia	A member of Coniferales, possibly a member of the stem group of Pinaceae; a species of Schizolepidopsis.
Stephanospermum shuichengensis[20]	Sp. nov.	Valid	Spencer et al.	Permian (Lopingian)	Wangjiazhai Formation	China	A member of Medullosales, a species of Stephanospermum.

Other plants

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Ashicaulis beipiaoensis[21]	Sp. nov	Valid	Tian et al.	Middle Jurassic	Tiaojishan Formation	China	A member of Osmundaceae, a species of Ashicaulis.
Bryiidites[22]	Gen. et sp. nov.	Valid	Barclay et al.	Late Cretaceous (late Cenomanian)	Middle Dakota Formation	United States	A member of Bryidae of uncertain phylogenetic placement. The type species is Bryiidites utahensis.
Franhueberia[23]	Gen. et sp. nov.	Valid	Hoffman & Tomescu	Devonian (Emsian)	Battery Point Formation	Canada	A basal member of Euphyllophytina. The type species is Franhueberia gerriennei.
Palaeosorum[24]	Gen. et sp. nov	Valid	Jacques & Zhou in Jacques, Su & Zhou	Miocene	Dajie Formation	China	A member of Polypodiaceae. The type species is Palaeosorum ellipticum.

Arthropods

Bryozoans

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Adlatipora[25]	Gen. et sp. nov	Valid	Gautier, Jackson & McKinney	Permian		United States	An acanthocladiid bryozoan. The type species is Adlatipora fossulata.
Basyaylella[26]	Gen. et sp. nov	Valid	Zágoršek & Gordon	Late Tortonian	Mut Basin	Turkey	An ascophoran bryozoan. Its type species is Basyaylella elsae.
Charixa goshouraensis[27]	Sp. nov	Valid[28]	Dick et al.	Cretaceous (late Albian or early Cenomanian)		Japan	A malacostegan, a species of Charixa.
Conopeum wilsoni[29]	Sp. nov	Valid	Di Martino & Taylor	Late Cretaceous (Late Campanian or Maastrichtian)	Simsima Formation	United Arab Emirates–Oman border region	A species of Conopeum.
Haplostoechios[27]	Gen. et 2 sp. nov	Valid[28]	Dick et al.	Cretaceous (late Albian or early Cenomanian)		Japan	A malacostegan. Genus contains two species: H. hayamiae and H. clusum.
Heloclema uralicum[30]	Sp. nov	Valid	Tolokonnikova	Early Carboniferous		Russia	A species of Heloclema.
Monoceratopora whybrowi[29]	Sp. nov	Valid	Di Martino & Taylor	Late Cretaceous (Late Campanian or Maastrichtian)	Simsima Formation	United Arab Emirates–Oman border region	A species of Monoceratopora.
Ostrovskia[26]	Gen. et sp. nov	Valid	Zágoršek & Gordon	Late Tortonian	Mut Basin	Turkey	An ascophoran bryozoan. Its type species is Ostrovskia triforamina.
Primorella kodinkensis[30]	Sp. nov	Valid	Tolokonnikova	Early Carboniferous		Russia	A species of Primorella.
Tyloporella smithi[29]	Sp. nov	Valid	Di Martino & Taylor	Late Cretaceous (Late Campanian or Maastrichtian)	Simsima Formation	United Arab Emirates–Oman border region	A species of Tyloporella.
Wilbertopora cheethami[29]	Sp. nov	Valid	Di Martino & Taylor	Late Cretaceous (Late Campanian or Maastrichtian)	Simsima Formation	United Arab Emirates–Oman border region	A species of Wilbertopora.

Brachiopods

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Arenorthis paranaensis[31]	Sp. nov	Valid	Benedetto, Halpern & Inchausti	Late Ordovician (Hirnantian)	Eusebio Ayala Formation	Paraguay	A species of Arenorthis.
Askepasma saproconcha[32]	sp nov	Valid	Topper et al..	Lower Cambrian	Wirrapowie Limestone	Australia	A member of Paterinata, a species of Askepasma.
Biseptata[33]	Gen. et sp. nov	Valid	Madison	Ordovician		Russia	A member of Strophomenida. The type species is Biseptata briani.
Menathyris[34]	Gen. et sp. nov	Valid	Feldman	Middle Triassic (early Ladinian)	Saharonim Formation	Israel	A zeilleriid terebratulid. The type species is Menathyris wilsoni.
Neochonetes (Huangichonetes) archboldi[35]	Sp. nov	Valid	Zhang et al.	Late Permian (Changhsingian)		China	A member of Rugosochonetidae, a species of Neochonetes.
Neochonetes semicircularis[35]	Sp. nov	Valid	Zhang et al.	Late Permian (Changhsingian)		China	A member of Rugosochonetidae, a species of Neochonetes.
Neochonetes (Sommeriella) rectangularis[35]	Sp. nov	Valid	Zhang et al.	Late Permian (Changhsingian)		China	A member of Rugosochonetidae, a species of Neochonetes.
Neochonetes (Sommeriella) waterhousei[35]	Sp. nov	Valid	Zhang et al.	Late Permian (Changhsingian)		China	A member of Rugosochonetidae, a species of Neochonetes.
Plectothyrella? itacurubiensis[31]	Sp. nov	Valid	Benedetto, Halpern & Inchausti	Late Ordovician (Hirnantian)	Eusebio Ayala Formation	Paraguay	Possibly a species of Plectothyrella.
Proclinorthis[36]	Gen. et sp. nov	Valid	Colmenar, Villas & Vizcaïno	Late Ordovician	Glauzy Formation	France	A platyorthid. The type species is Proclinorthis vailhanensis.
Pseudocrania insperata[37]	Sp. nov	Valid	Bassett et al.	Middle Ordovician (Darriwilian)	Lashkarak Formation	Iran	A member of Craniida, a species of Pseudocrania.
Sphenope[38]	Gen. et comb. et sp. nov	Valid	Vörös	Late Jurassic to Early Cretaceous (Tithonian to Berriasian)		Hungary  Italy	A pygopid terebratulid. A new genus for "Terebratula" misilmerensis Gemmellaro (1871); genus also contains the new species Sphenope bifida.
Tethyochonetes cheni[35]	Sp. nov	Valid	Zhang et al.	Late Permian (Changhsingian)		China	A member of Rugosochonetidae, a species of Tethyochonetes.
Tethyochonetes sheni[35]	Sp. nov	Valid	Zhang et al.	Late Permian (Changhsingian)		China	A member of Rugosochonetidae, a species of Tethyochonetes.

Molluscs

Echinoderms

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Abertella miskellyi[39]	Sp. nov	Valid	Kroh et al.	Miocene	Camarones Formation	Argentina	An abertellid sand dollar, a species of Abertella.
Alternacantha[40]	Gen. et sp. nov	Valid	Thuy & Meyer	Middle Jurassic		Switzerland	An ophiacanthid brittle star. The type species is Alternacantha occulta.
Aragocystites[41]	Gen. et sp. nov	Valid	Zamora	Middle Cambrian	Murero Formation	Spain	An edrioasteroid. The type species is Aragocystites belli.
Bathyovulaster[42]	Gen. et sp. nov	Valid	Smith	Santonian		Italy	A spatangoid. The type species is Bathyovulaster disjunctus.
Bellastrella[43]	Gen. et sp. nov	Valid	Müller, Hahn and Bohatý	Middle Devonian		Germany	An agelacrinitid edrioasteroid. The type species is Bellastrella eifeliana.
Ctenocrinus branisai[44]	Sp. nov	Valid	Thompson, Ausich & Smith	Early Devonian (Emsian)	Belèn Formation	Bolivia	A crinoid, a species of Ctenocrinus.
Cyclolampas altus[45]	Sp. nov	Valid	Saucede et al.	Middle Jurassic (late Callovian)		France	An echinoid, a species of Cyclolampas.
Eozonella[46]	Gen. et sp. nov	Valid	Thuy, Marty & Comment	Late Jurassic (late Oxfordian)		Switzerland	An ophiolepidid brittle star. The type species is Eozonella bergeri.
Graciacystis[47]	Gen. et sp. nov	Valid	Zamora, Rahman & Smith	Middle Cambrian		Spain	A basal cinctan echinoderm. The type species is Graciacystis ambigua.
Griphocrinus pirovanoi[44]	Sp. nov	Valid	Thompson, Ausich & Smith	Early Devonian (Emsian)	Icla Formation	Bolivia	A crinoid, a species of Griphocrinus.
Hanshessia[40]	Gen. et sp. nov	Valid	Thuy & Meyer	Middle Jurassic		Switzerland	An ophiacanthid brittle star. The type species is Hanshessia trochitophila.
Juracantha[46]	Gen. et sp. nov	Valid	Thuy, Marty & Comment	Late Jurassic (late Oxfordian)		Switzerland	An ophiacanthid brittle star. The type species is Juracantha hottingeri.
Kosachenkoastrus[48]	Gen. et sp. nov	Valid	Rozhnov	Middle Ordovician		Russia	A member of Parablastoidea. The type species is Kosachenkoastrus volkhovensis.
Lutocrinus[44]	Gen. et sp. nov	Valid	Thompson, Ausich & Smith	Early Devonian (Emsian)	Icla Formation	Bolivia	A crinoid. The type species is Lutocrinus boliviaensis.
Ophiotholia aurora[40]	Sp. nov	Valid	Thuy & Meyer	Middle Jurassic		Switzerland	An ophiomycetid brittle star, a species of Ophiotholia.
Rhenopyrgus piojoensis[49]	Sp nov	Valid	Sumrall, Heredia, Rodríguez & Mestre	Silurian (lower Ludlow)	Los Espejos Formation	Argentina	A rhenopyrgid edrioasteroid, a species of Rhenopyrgus.
Simakocrinus[50]	Gen. et 2 sp. nov.	Valid	Prokop	Devonian (Pragian, Emsian and Eifelian)		Czech Republic	A crinoid belonging to the group Disparida, possibly a member of the family Myelodactylidae. The type species is Simakocrinus facilis sp. nov (col.); genus also contains Simakocrinus diligens sp. nov (col.).

Conodonts

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Acanthodus humachensis[51]	Sp nov	Valid	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Acanthodus.
Acanthodus raqueli[51]	Sp nov	Valid	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Acanthodus.
Acodus primitivus[51]	Sp nov	Valid	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Acodus.
Acuminatella[52]	Gen. et 2 sp. nov	Valid	Orchard	Late Triassic (late Carnian to early Norian)	Pardonet Formation	Canada	A gondolellid ozarkodinid. The type species is Acuminatella acuminata; genus also contains Acuminatella angusta.
Bispathodus ultimus bartzschi[53]	Subsp nov	Valid	Kononova & Weyer	Devonian (late Famennian)		Germany	A subspecies of Bispathodus ultimus.
Carnepigondolella eozoae[52]	Sp. nov	Valid	Orchard	Late Triassic (late Carnian)	Ludington Formation	Canada	A gondolellid ozarkodinid, a species of Carnepigondolella.
Carnepigondolella medioconstricta[52]	Sp. nov	Valid	Orchard	Late Triassic (late Carnian)	Ludington Formation	Canada	A gondolellid ozarkodinid, a species of Carnepigondolella.
Furnishina leei[54]	Sp. nov	Valid	Lee	Cambrian	Machari Formation	South Korea
Histiodella wuhaiensis[55]	Sp. nov	Valid	Wang et al.	Ordovician	Lower Klimoli Formation	China
Idiognathodus craticulinodosus[56]	Sp. nov	Valid	Ishida, Suzuki & Inada	Carboniferous (middle–late Bashkirian to early Moscovian)	Ko-yama Limestone Group	Japan
Kallidontus gondwanicus[51]	Sp nov	Valid	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Kallidontus.
Kraussodontus[52]	Gen. et sp. nov	Valid	Orchard	Late Triassic (late Carnian)	Pardonet Formation	Canada	A gondolellid ozarkodinid. The type species is Kraussodontus peteri.
Mesogondolella daheshenensis[57]	Sp nov	Valid	Wang in Zhou et al.	Permian (probably Wordian)	Daheshen Formation	China	A species of Mesogondolella.
Mesogondolella subgracilis[57]	Sp nov	Valid	Wang in Zhou et al.	Permian (probably Wordian)	Daheshen Formation	China	A species of Mesogondolella.
Muellerilepis[58]	Nom. nov	Valid	Bardashev & Bardasheva	Middle Devonian (Givetian)		Tajikistan	A replacement name for Muellerina Bardashev & Bardasheva (2012) (preoccupied).
Naimanodus[59]	Gen. et sp. nov	Valid	Tolmacheva	Middle Ordovician		Kazakhstan  Kyrgyzstan  Russia	The type species is Naimanodus degtyarevi.
Neopolygnathus margaretae[53]	Sp nov	Valid	Kononova & Weyer	Devonian (late Famennian)		Germany	A species of Neopolygnathus.
Notiodella[60]	Gen. et sp. nov	Valid	Aldridge et al.	Late Ordovician	Soom Shale Member, Cedarberg Formation	South Africa	A member of Prioniodontida belonging to the family Balognathidae. The type species is Notiodella keblon.
Palmatolepis khaensis[61]	Sp. nov	Valid	Savage	Late Devonian		Thailand
Palmatolepis marki[61]	Sp. nov	Valid	Savage	Late Devonian		Thailand
Palmatolepis subperlobata lapoensis[61]	Subsp. nov	Valid	Savage	Late Devonian		Thailand
Parapetella[52]	Gen. et sp. nov	Valid	Orchard	Late Triassic (late Carnian to early Norian)	Pardonet Formation	Canada	A gondolellid ozarkodinid. The type species is Parapetella prominens.
Polygnathus burretti[61]	Sp. nov	Valid	Savage	Late Devonian		Thailand
Polygnathus communis hanensis[61]	Subsp. nov	Valid	Savage	Late Devonian		Thailand
Polygnathus communis namdipensis[61]	Subsp. nov	Valid	Savage	Late Devonian		Thailand
Polygnathus communis phaphaensis[61]	Subsp. nov	Valid	Savage	Late Devonian		Thailand
Polygnathus crassulus salapensis[61]	Subsp. nov	Valid	Savage	Late Devonian		Thailand
Polygnathus extralobatus phoensis[61]	Subsp. nov	Valid	Savage	Late Devonian		Thailand
Polygnathus hojedki[62]	Sp. nov	Valid	Gholamalian et al.	Devonian		Iran	A species of Polygnathus.
Polygnathus pseudocostatus[63]	Sp nov	Valid	Klapper & Vodrážková	Devonian		United States	A species of Polygnathus.
Primatella[52]	Gen. et comb. et 2 sp. nov	Valid	Orchard	Late Triassic (late Carnian to early Norian)	Pardonet Formation	Canada	A gondolellid ozarkodinid. A new genus for "Epigondolella" primitia Mosher (1970); genus also contains new species Primatella asymmetrica and Primatella conservativa.
Pseudopolygnathus granulosus laepensis[61]	Subsp. nov	Valid	Savage	Late Devonian		Thailand
Pseudopolygnathus granulosus salawinensis[61]	Subsp. nov	Valid	Savage	Late Devonian		Thailand
Pseudopolygnathus maepoensis[61]	Sp. nov	Valid	Savage	Late Devonian		Thailand
Quadralella[52]	Gen. et sp. nov	Valid	Orchard	Late Triassic (late Carnian)	Ludington Formation	Canada	A gondolellid ozarkodinid. The type species is Quadralella lobata.
Siphonodella banraiensis[61]	Sp. nov	Valid	Savage	Late Devonian		Thailand
Tilcarodus[51]	Gen. et comb. nov	Valid	Zeballo & Albanesi	Early Ordovician	Rupasca Formation	Argentina	A new genus for "Utahconus" humahuacensis Albanesi and Aceñolaza (2005).
Trapezognathus? primitivus[64]	Sp. nov	Valid	Voldman, Albanesi & Zeballo in Voldman et al.	Early Ordovician	Acoite Formation	Argentina	Originally described as a possible species of Trapezognathus; subsequently made the type species of a separate genus Zentagnathus.[65]
Utahconus purmamarcensis[51]	Sp nov	Valid	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Utahconus.
Utahconus scandodiformis[51]	Sp nov	Valid	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Utahconus.
Utahconus tortibasis[51]	Sp nov	Valid	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Utahconus.
Variabiloconus crassus[51]	Sp nov	Valid	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Variabiloconus.

Fishes

Amphibians

Newly named temnospondyls

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Broiliellus reiszi[66]	Sp. nov	Valid	Holmes, Berman & Anderson	Early Permian	El Cobre Canyon Formation	United States	A dissorophid, a species of Broiliellus.
Parotosuchus ptaszynskii[67]	Sp nov	Valid	Sulej & Niedźwiedzki	Late Olenekian		Poland	A species of Parotosuchus.
Reiszerpeton[68]	Gen. et sp. nov	Valid	Maddin et al.	Early Permian	Archer City Formation	United States	A dissorophid. The type species is Reiszerpeton renascentis.
Scapanops[69]	Gen. et sp. nov	Valid	Schoch & Sues	Early Permian	Nocona Formation	United States	A dissorophid. The type species is Scapanops neglecta.
Tersomius dolesensis[70]	Sp nov	Valid	Anderson & Bolt	Early Permian		United States	A species of Tersomius.
Trimerorhachis greggi[71]	Sp nov	Valid	Milner & Schoch	Early Permian		United States	A species of Trimerorhachis.

Newly named lepospondyls

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Batropetes niederkirchensis[72]	Sp. nov	Valid	Glienke	Early Permian (Rotliegend)	Saar–Nahe Basin	Germany	A species of Batropetes.
Huskerpeton[73]	Gen. et sp. nov	Valid	Huttenlocker et al.	Early Permian	Eskridge Formation	United States	A recumbirostran. The type species is Huskerpeton englehorni.
Proxilodon[73]	Gen. et comb. nov	Valid	Huttenlocker et al.	Early Permian	Speiser Formation	United States	A recumbirostran, a new genus for "Euryodus" bonneri.

Newly named lissamphibians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Aerugoamnis[74]	Gen. et sp. nov	Valid	Henrici, Báez & Grande	Early Eocene (Wasatchian)	Green River Formation	United States	A relative of parsley frogs. The type species is Aerugoamnis paulus.
Eobarbourula[75]	Gen. et sp. nov	Valid	Folie et al..	Early Eocene	Cambay Shale Formation	India	A bombinatorid. Its type species is Eobarbourula delfinoi.
Gracilibatrachus[76]	Gen. et sp. nov	Valid	Báez	Early Cretaceous (late Barremian)		Spain	A relative of the family Pipidae. The type species is Gracilibatrachus avallei.
Iberobatrachus[76]	Gen. et sp. nov	Valid	Báez	Early Cretaceous (late Barremian)		Spain	A close relative of the genus Discoglossus. The type species is Iberobatrachus angelae.
Ichthyosaura randeckensis[77]	Sp. nov	Valid	Schoch & Rasser	Miocene		Germany	A newt related to the Alpine newt.
Indorana[75]	Gen. et sp. nov	Valid	Folie et al..	Early Eocene	Cambay Shale Formation	India	A possible rhacophorid. Its type species is Indorana prasadi.
Leiopelma acricarina[78]	Sp. nov	Valid	Worthy et al.	Early Miocene		New Zealand	A member of Leiopelmatidae.
Leiopelma miocaenale[78]	Sp. nov	Valid	Worthy et al.	Early Miocene		New Zealand	A member of Leiopelmatidae.
Liaobatrachus zhaoi[79]	Sp. nov	Valid	Dong et al..	Early Cretaceous (Barremian)	Yixian Formation	China	An anuran of uncertain phylogenetic placement, a species of Liaobatrachus.
Paranecturus[80]	Gen. et sp. nov	Valid	Demar	Late Cretaceous (Maastrichtian)	Hell Creek Formation	United States	A mudpuppy. The type species is Paranecturus garbanii.
Ukrainurus[81]	Gen. et sp. nov	Valid	Vasilyan et al.	Miocene		Ukraine	A relative of cryptobranchids. The type species is Ukrainurus hypsognathus.
Wesserpeton[82]	Gen. et sp. nov	Valid	Sweetman & Gardner	Barremian	Wessex Formation	England	An albanerpetontid.

Turtles

Newly named turtles

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Adocus inexpectatus[83]	Sp. nov	Valid	Danilov et al.	Late Eocene	Youganwo Formation	China	An adocid, a species of Adocus.
Atoposemys[84]	Gen. et sp. nov	Valid	Hutchison	Early Paleocene (Puercan)	Fort Union Formation	United States	A trionychid. The type species is Atoposemys entopteros.
Bairdemys healeyorum[85]	Sp. nov	Valid	Weems & Knight	Late Oligocene	Chandler Bridge Formation	United States	A podocnemidid pleurodiran, a species of Bairdemys.
Basilemys gaffneyi[86]	Sp. nov	Valid	Sullivan, Jasinski & Lucas	Late Campanian		United States	A nanhsiungchelyid turtle, a species of Basilemys.
Brachyopsemys[87]	Gen. et sp. nov	Valid	Tong & Meylan	Early Paleocene	Ouled Abdoun Basin	Morocco	A cryptodiran closely related to Sandownia. The type species is Brachyopsemys tingitana.
Camerochelys[88]	Gen. et sp. nov	Valid	Pérez-García & Murelaga	Early Cretaceous (Hauterivian or Barremian)	Cameros Basin	Spain	A member of Pan-Cryptodira (the clade containing living cryptodirans and all turtles sharing a more recent common ancestor with them than with pleurodirans), possibly a xinjiangchelyid. The type species is Camerochelys vilanovai.
Cardichelyon[84]	Gen. et sp. nov	Valid	Hutchison	Early Eocene (Wasatchian)		United States	A platysternid. The type species is Cardichelyon rogerwoodi.
?Centrochelys marocana[89]	Sp. nov	Valid	Gmira et al.	Pliocene		Morocco	A tortoise, possibly a species of Centrochelys.
Changmachelys[90]	Gen. et sp. nov	Valid	Brinkman et al.	Early Cretaceous	Xiagou Formation	China	A member of (likely paraphyletic) group "Macrobaenidae". The type species is Changmachelys bohlini.
Chelodina (Chelodina) murrayi[91]	Sp. nov	Valid	Yates	Late Miocene	Waite Formation	Australia	A chelid, a species of Chelodina.
Chelonoidis lutzae[92]	Sp. nov	Valid	Zacarías et al.	Late Pleistocene (58–22 ka)		Argentina	A tortoise, a species of Chelonoidis.
Chrysemys isoni[93]	Sp. nov	Valid	Weems & George	Miocene	Calvert Formation	United States	A relative of the painted turtle.
Cuora chiangmuanensis[94]	Sp. nov	Valid	Naksri et al.	Late Middle or early Late Miocene		Thailand	An Asian box turtle.
Floridemys hurdi[93]	Sp. nov	Valid	Weems & George	Miocene	Calvert Formation	United States	A tortoise, a species of Floridemys.
Kappachelys[95]	Gen. et sp. nov	Valid	Hirayama, Isaji & Hibino	Early Cretaceous	Akaiwa Formation	Japan	A relative of trionychids. The type species is Kappachelys okurai.
Laganemys[96]	Gen. et sp. nov	Valid	Sereno & ElShafie	Aptian or Albian	Elrhaz Formation	Niger	An araripemydid pleurodiran. The type species is Laganemys tenerensis.
Neochelys liriae[97]	Sp. nov	Valid	Pérez-García & de Lapparent de Broin	Eocene (Ypresian)		France	A podocnemidid, a species of Neochelys.
Neurankylus lithographicus[98]	Sp. nov	Valid	Larson et al.	Santonian	Milk River Formation	Canada	A neurankyline baenid, a species of Neurankylus.
Nostimochelone[99]	Gen. et sp. nov	Valid	Georgalis et al.	Early Miocene	Zeugostasion Formation	Greece	A pleurodiran, a member of Podocnemidoidea. The type species is Nostimochelone lampra.
Ocadia tanegashimensis[100]	Sp. nov	Valid	Takahashi et al.	Early middle Miocene	Kawachi Formation	Japan	A geoemydid, a species of Ocadia.
Ocepechelon[101]	Gen. et sp. nov	Valid	Bardet et al.	Late Cretaceous (late Maastrichtian)	Oulad Abdoun Basin	Morocco	A relative of dermochelyids and protostegids. The type species is Ocepechelon bouyai.
Paramongolemys[102]	Gen. et sp. nov	Valid	Danilov & Sukhanov	Late Paleocene	Naranbulak Formation	Mongolia	A basal member of Testudinoidea. The type species is Paramongolemys khosatzkyi.
Planetochelys dithyros[84]	Sp. nov	Valid	Hutchison	Early Eocene (Wasatchian)		United States	A relative of trionychids, a species of Planetochelys.
Psilosemys[84]	Gen. et sp. nov	Valid	Hutchison	Early Eocene (Wasatchian)		United States	The earliest North American emydid. The type species is Psilosemys wyomingensis.
Scabremys[86]	Gen. et comb. nov	Valid	Sullivan, Jasinski & Lucas	Late Campanian		United States	A baenid cryptodiran, a new genus for "Baena" ornata Gilmore (1935).
Shandongemys[103]	Gen. et sp. nov	Valid	Li et al.	Late Cretaceous		China	A lindholmemydid testudinoid. The type species is Shandongemys dongwuica.
Sinemys brevispinus[104]	Sp. nov	Valid	Tong & Brinkman	Early Cretaceous	Luohandong Formation	China	A sinemydid cryptodiran, a species of Sinemys.
Spoochelys[105]	Gen. et sp. nov	Valid	Smith & Kear	Early or middle Albian	Griman Creek Formation	Australia	A meiolaniid-like turtle. The type species is Spoochelys ormondea.
Testudo oughlamensis[89]	Sp. nov	Valid	Gmira et al.	Pliocene		Morocco	A tortoise, a species of Testudo.
Trapalcochelys[106]	Gen. et sp. nov	Valid	Sterli, de la Fuente & Cerda	Late Cretaceous (late Campanian to early Maastrichtian)	Allen Formation	Argentina	A relative of meiolaniids. The type species is Trapalcochelys sulcata.
Tullochelys[84]	Gen. et sp. nov	Valid	Hutchison	Maastrichtian to early Paleocene (Puercan)	Fort Union Formation Hell Creek Formation	United States	A chelydrid. The type species is Tullochelys montanus.
Xenochelys floridensis[107]	Sp. nov	Valid	Bourque	Late Oligocene (early Arikareean)		United States	A kinosternid, a species of Xenochelys.
Xinjiangchelys radiplicatoides[108]	Sp. nov	Valid	Brinkman et al.	Late Middle or early Late Jurassic	Shishugou Formation	China	A species of Xinjiangchelys.
Xinjiangchelys wusu[109]	Sp. nov	Valid	Rabi et al.	Middle Jurassic	?Qigu Formation	China	A species of Xinjiangchelys.

Thalattosaurs

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Concavispina[110]	Gen. et sp. nov	Valid	Zhao et al.	Triassic	Xiaowa Formation	China	A thalattosaurid thalattosaur. The type species is Concavispina biseridens

Ichthyopterygians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Chaohusaurus zhangjiawanensis[111]	Sp. nov	Valid	Chen et al.	Early Triassic	Jialingjiang Formation	China	A species of Chaohusaurus.
Gulosaurus[112]	Gen. et sp. nov	Valid	Cuthbertson, Russell & Anderson	Early Triassic	Sulphur Mountain Formation	Canada	A grippidian. The type species is Gulosaurus helmi.
Malawania[113]	Gen. et sp. nov	Valid	Fischer et al.	Early Cretaceous (late Hauterivian to Barremian)		Iraq	A basal member of Thunnosauria. The type species is Malawania anachronus.
Thalattoarchon[114]	Gen. et sp. nov	Valid	Fröbisch et al.	Middle Triassic		United States	A merriamosaur ichthyosaur. The type species is Thalattoarchon saurophagis

Lepidosauromorphs

Newly named sauropterygians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Cryonectes[115]	Gen. et sp. nov	Valid	Vincent, Bardet & Mattioli	Pliensbachian		France	A pliosaurid. The type species is Cryonectes neustriacus
Gronausaurus[116]	Gen. et sp. nov	Valid	Hampe	Early Cretaceous (Berriasian)		Germany	A leptocleidid[116] or an elasmosaurid.[117] The type species is Gronausaurus wegneri
Megacephalosaurus[118]	Gen. et sp. nov	Valid	Schumacher, Carpenter & Everhart	Late Cretaceous (middle Turonian)		United States	A pliosaurid. The type species is Megacephalosaurus eulerti.
Palatodonta[119]	Gen. et sp. nov	Valid	Neenan, Klein & Scheyer	Early Middle Triassic		Netherlands	A relative of placodonts. The type species is Palatodonta bleekeri.
Pliosaurus carpenteri[117]	Sp. nov	Valid	Benson et al.	Late Jurassic	Kimmeridge Clay Formation	United Kingdom	A pliosaurid, a species of Pliosaurus
Pliosaurus kevani[117]	Sp. nov	Valid	Benson et al.	Late Jurassic	Kimmeridge Clay Formation	United Kingdom	A pliosaurid, a species of Pliosaurus
Pliosaurus westburyensis[117]	Sp. nov	Valid	Benson et al.	Late Jurassic	Kimmeridge Clay Formation	United Kingdom	A pliosaurid, a species of Pliosaurus
Serpianosaurus germanicus[120]	Sp. nov	Valid	Diedrich	Middle Triassic (Anisian)		Germany	A pachypleurosaur, a species of Serpianosaurus.

Newly named rhynchocephalians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Sphenotitan[121]	Gen. et sp. nov	Valid	Martínez et al.	Late Triassic	Quebrada del Barro Formation	Argentina	An opisthodontian rhynchocephalian. The type species is Sphenotitan leyesi.

Newly named lizards

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Apsgnathus[122]	Gen. et sp. nov	Valid	Nydam, Rowe & Cifelli	Late Cretaceous (late Campanian, Judithian)	Aguja Formation	United States	A member of Scincomorpha of uncertain phylogenetic placement. The type species is Apsgnathus triptodon.
Arcanosaurus[123]	Gen. et sp. nov	Valid	Houssaye et al.	Early Cretaceous (Barremian or Aptian)	Cameros Basin	Spain	An anguimorph lizard, possibly a varanoid. The type species is Arcanosaurus ibericus.
Arpadosaurus sepulchralis[124]	Sp. nov	Valid	Smith & Gauthier	Early Eocene	Wasatch Formation	United States	A member of Glyptosaurinae, a species of Arpadosaurus.
Barbaturex[125]	Gen. et sp. nov	Valid	Head et al.	Late middle Eocene (37.2 ± 1.3 Ma)	Pondaung Formation	Myanmar	An iguanian lizard related to the genus Uromastyx. The type species is Barbaturex morrisoni.
Bicuspidon smikros[126]	Sp. nov	Valid	Nydam	Late Cretaceous (Cenomanian)	Dakota Formation	United States	A member of Polyglyphanodontinae, a species of Bicuspidon.
Blanosaurus[127]	Gen. et sp. nov	Valid	Folie, Smith & Smith	Early Eocene		Belgium  France	An amphisbaenian. The type species is Blanosaurus primeocaenus.
Camptognathosaurus[127]	Gen. et sp. nov	Valid	Folie, Smith & Smith	Late Paleocene		France	An amphisbaenian. The type species is Camptognathosaurus parisiensis.
Catactegenys[122]	Gen. et sp. nov	Valid	Nydam, Rowe & Cifelli	Late Cretaceous (late Campanian, Judithian)	Aguja Formation	United States	A night lizard. The type species is Catactegenys solaster.
Cemeterius[128][129]	Gen. et sp. nov	Valid	Longrich, Bhullar & Gauthier	Late Cretaceous (late Maastrichtian)	Lance Formation	United States	A platynotan lizard of uncertain phylogenetic placement. The type species is Cemeterius monstrosus.
Cnodontosaurus[126]	Gen. et sp. nov	Valid	Nydam	Late Cretaceous (Cenomanian)	Dakota Formation	United States	A member of Anguimorpha related to xenosaurids. The type species is Cnodontosaurus suchockii.
Dakotasaurus[126]	Gen. et sp. nov	Junior homonym	Nydam	Late Cretaceous (Cenomanian)	Dakota Formation	United States	A member of Scincomorpha of uncertain phylogenetic placement, assigned by Nydam (2013) to an informal paramacellodid-cordylid grade.[126] The type species is Dakotasaurus gillettorum. The generic name turned out to be preoccupied by Dakotasaurus Branson & Mehl (1932); the lizard genus was subsequently renamed Dakotaseps.[130]
Desertiguana[131]	Gen. et sp. nov	Valid	Alifanov	Late Cretaceous		Mongolia	A phrynosomatid lizard. The type species is Desertiguana gobiensis.
Distortodon[132]	Gen. et sp. nov	Valid	Makádi	Late Cretaceous (Santonian)		Hungary	A member of Polyglyphanodontinae. The type species is Distortodon rhomboideus.
Entomophontes[124]	Gen. et 2 sp. nov	Valid	Smith & Gauthier	Early Eocene	Wasatch Formation	United States	A relative of species belonging to the genus Xenosaurus. Genus contains two species: Entomophontes incrustatus and Entomophontes hutchisoni.
Eonatator coellensis[133]	Sp. nov	Valid	Páramo	Campanian		Colombia	A mosasaur, a new species of Eonatator.
Glyptosaurus agmodon[124]	Sp. nov	Valid	Smith & Gauthier	Early Eocene	Wasatch Formation	United States	A member of Glyptosaurinae, a species of Glyptosaurus.
Heterodontagama[134]	Gen. et sp. nov	Valid	Rana et al.	Early Eocene (early to middle Ypresian)	Cambay Shale Formation	India	A member of Priscagamidae. The type species is Heterodontagama borsukae.
Indiagama[134]	Gen. et sp. nov	Valid	Rana et al.	Early Eocene (early to middle Ypresian)	Cambay Shale Formation	India	A member of Agamidae. The type species is Indiagama gujarata.
Lamiasaura[128][129]	Gen. et sp. nov	Valid	Longrich, Bhullar & Gauthier	Late Cretaceous (late Maastrichtian)	Lance Formation	United States	A squamate of uncertain phylogenetic placement. The type species is Lamiasaura ferox.
Lonchisaurus[128][129]	Gen. et sp. nov	Valid	Longrich, Bhullar & Gauthier	Late Cretaceous (late Maastrichtian)	Lance Formation	United States	A scincomorph lizard of uncertain phylogenetic placement. The type species is Lonchisaurus trichurus.
Monocnemodon[126]	Gen. et sp. nov	Valid	Nydam	Late Cretaceous (Santonian)	Straight Cliffs Formation	United States	A member of Scincomorpha of uncertain phylogenetic placement, assigned by Nydam (2013) to an informal paramacellodid-cordylid grade.[126] The type species is Monocnemodon syphakos.
Obamadon[128][129]	Gen. et sp. nov	Valid	Longrich, Bhullar & Gauthier	Late Cretaceous (late Maastrichtian)	Hell Creek Formation Lance Formation	United States	A polyglyphanodontian lizard of uncertain phylogenetic placement. The type species is Obamadon gracilis.
Odaxosaurus roosevelti[126]	Sp. nov	Valid	Nydam	Late Cretaceous	Kaiparowits Formation	United States	A member of Anguidae, a species of Odaxosaurus.
Palaeoxantusia amara[124]	Sp. nov	Valid	Smith & Gauthier	Early Eocene	Wasatch Formation	United States	A night lizard, a species of Palaeoxantusia.
Palepidophyma[124]	Gen. et 2 sp. nov	Valid	Smith & Gauthier	Early Eocene	Wasatch Formation	United States	A night lizard. Genus contains two species: Palepidophyma paradisa and Palepidophyma lilliputiana.
Parasaniwa cynochoros[126]	Sp. nov	Valid	Nydam	Late Cretaceous	Kaiparowits Formation	United States	A member of Platynota of uncertain phylogenetic placement, a species of Parasaniwa.
Pariguana[128][129]	Gen. et sp. nov	Valid	Longrich, Bhullar & Gauthier	Late Cretaceous (late Maastrichtian)	Lance Formation	United States	An iguanid lizard of uncertain phylogenetic placement. The type species is Pariguana lancensis.
Pelsochamops[135]	Gen. et sp. nov	Valid	Makádi	Late Cretaceous (Santonian)	Csehbánya Formation	Hungary	A chamopsiid. The type species is Pelsochamops infrequens.
Plesiolacerta eratosthenesi[136]	Sp. nov	Valid	Čerňanský & Augé	Late Oligocene		Germany	A lacertid lizard, a species of Plesiolacerta.
Polyodontobaena[127]	Gen. et sp. nov	Valid	Folie, Smith & Smith	Early Paleocene	Hainin Formation	Belgium	An amphisbaenian. The type species is Polyodontobaena belgica.
Provaranosaurus fatuus[124]	Sp. nov	Valid	Smith & Gauthier	Early Eocene	Wasatch Formation	United States	A relative of the Chinese crocodile lizard, a species of Provaranosaurus.
Romeosaurus[137]	Gen. et 2 sp. nov	Valid	Palci, Caldwell & Papazzoni	Late Cretaceous (early Turonian to early Santonian)		Italy	A mosasaur related to Russellosaurus and Yaguarasaurus. Genus contains two species: Romeosaurus fumanensis and Romeosaurus sorbinii.
Schillerosaurus[138]	Nom. nov	Valid	Nydam, Chure & Evans	Late Jurassic	Morrison Formation	United States	A replacement name for Schilleria Evans & Chure (1999).
Scincoideus grassator[124]	Sp. nov	Valid	Smith & Gauthier	Early Eocene	Wasatch Formation	United States	A member of Lacertiformes, a species of Scincoideus.
Socognathus brachyodon[128][129]	Sp. nov	Valid	Longrich, Bhullar & Gauthier	Late Cretaceous (late Maastrichtian)	Lance Formation	United States	A chamopsiid polyglyphanodontian lizard, a species of Socognathus.
Suratagama[134]	Gen. et sp. nov	Valid	Rana et al.	Early Eocene (early to middle Ypresian)	Cambay Shale Formation	India	A member of Agamidae. The type species is Suratagama neeraae.
Suzanniwana revenanta[124]	Sp. nov	Valid	Smith & Gauthier	Early Eocene	Wasatch Formation	United States	A member of Pleurodonta, a species of Suzanniwana.
Tiliqua laticephala[139]	Sp. nov	Valid	Čerňanský & Hutchinson	Pliocene		Australia	A blue-tongued skink.
Utahgenys antongai[126]	Sp. nov	Valid	Nydam	Late Cretaceous (Cenomanian)	Dakota Formation	United States	A contogeniid scincomorph, a species of Utahgenys.
Webbsaurus[126]	Gen. et sp. nov	Valid	Nydam	Late Cretaceous (Cenomanian)	Dakota Formation	United States	A scincomorph related to skinks. The type species is Webbsaurus lofgreni.
Xestops savagei[124]	Sp. nov	Valid	Smith & Gauthier	Early Eocene	Wasatch Formation	United States	A member of Glyptosaurinae, a species of Xestops.

Newly named snakes

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Cerberophis[128][129]	Gen. et sp. nov	Valid	Longrich, Bhullar & Gauthier	Late Cretaceous (late Maastrichtian)	Hell Creek Formation	United States	An alethinophidian snake of uncertain phylogenetic placement. The type species is Cerberophis robustus.
Kataria[140]	Gen. et sp. nov	Valid	Scanferla et al.	Early Paleocene (Danian)	Santa Lucía Formation	Bolivia	A macrostomatan snake related to caenophidians. The type species is Kataria anisodonta.
Nidophis[141]	Gen. et sp. nov	Valid	Vasile, Csiki-Sava & Venczel	Late Cretaceous (Maastrichtian)	Haţeg Basin	Romania	A madtsoiid. The type species is Nidophis insularis.

Archosauromorphs

Newly named basal archosauromorphs

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Ankylosuchus[142]	Gen. et sp. nov	Valid	Lucas, Spielmann & Hunt	Late Triassic	Colorado City Formation	United States	A doswelliid. The type species is Ankylosuchus chinlegroupensis.
Asperoris[143]	Gen. et sp. nov	Valid	Nesbitt, Butler & Gower	Middle Triassic (probably late Anisian)	Manda beds	Tanzania	A non-archosaurian archosauriform of uncertain phylogenetic placement. The type species is Asperoris mnyama.
Fuyuansaurus[144]	Gen. et sp. nov	Valid	Fraser, Rieppel & Chun	Middle Triassic (Ladinian)	Falang Formation	China	A protorosaur. The type species is Fuyuansaurus acutirostris.
Jaxtasuchus[145]	Gen. et sp. nov	Valid[146]	Schoch & Sues	Middle Triassic (Ladinian)	Erfurt Formation	Germany	A doswelliid. The type species is Jaxtasuchus salomoni.
Machaeroprosopus lottorum[147]	Sp. nov	Valid	Hungerbühler et al.	Late Triassic	Cooper Canyon Formation	United States	A phytosaur, a species of Machaeroprosopus.
Macrocnemus obristi[148]	Sp. nov	Valid	Fraser & Furrer	Middle Triassic (Ladinian)	Prosanto Formation	Switzerland	A protorosaur, a species of Macrocnemus.
Wannia[149]	Gen. et comb. nov	Valid	Stocker	Late Triassic		United States	A phytosaur; a new genus for "Paleorhinus" scurriensis Langston (1949).

Archosaurs

Other reptiles

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Dolerosaurus[150]	Gen. et comb. nov	Valid	Butler	Late Triassic (late Carnian)	Lunz Formation	Austria	A diapsid of uncertain phylogenetic placement, a new genus for "Francosuchus" trauthi Huene (1939).
Ruhuhuaria[151]	Gen. et sp. nov	Valid	Tsuji, Sobral & Müller	Middle Triassic	Manda Formation	Tanzania	A procolophonoid parareptile, possibly an owenettid. The type species is Ruhuhuaria reiszi.

Synapsids

Non-mammalian synapsids

Research

• The postcranial skeleton of therocephalian Ictidosuchoides is described by Heidi Fourie (2013).^[152]

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Diegocanis[153]	Gen. et sp. nov	Valid	Martínez, Fernandez & Alcober	Late Triassic	Ischigualasto Formation	Argentina	A member of Eucynodontia closely related to Ecteninion and Trucidocynodon. The type species is Diegocanis elegans.
Mandagomphodon[154]	Gen. et comb. nov	Valid[155]	Hopson	Middle Triassic	Manda Formation	Tanzania	A traversodontid cynodont, a new genus for "Scalenodon" hirschsoni Crompton (1972).
Niassodon[156]	Gen. et sp. nov	Valid	Castanhinha et al.	Late Permian		Mozambique	A kingoriid emydopoid dicynodont. The type species is Niassodon mfumukasi.
Tambacarnifex[157]	Gen. et sp. nov	Valid[155]	Berman et al.	Early Permian	Tambach Formation	Germany	A varanodontine varanopid. The type species is Tambacarnifex unguifalcatus.

Mammals

Other animals

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Anomalocaris kunmingensis[158]	Sp. nov	Valid	Wang, Huang & Hu	Cambrian	Wulongqing Formation	China	A member of Anomalocarididae (a group of animals with uncertain phylogenetic placement, possibly stem-arthropods), a species of Anomalocaris.
Aulastraea crassicalix[159]	Sp. nov	Valid	Löser, Castro & Nieto	Early Cretaceous (Albian)		Spain	A stony coral, a species of Aulastraea.
Cambrocornulitus[160]	Gen. et sp. nov	Valid	Yang et al.	Early Cambrian	Qiongzhusi Formation	China	An animal of uncertain phylogenetic placement, possibly a member of Cornulitida. The type species is Cambrocornulitus rarus.
Conularia porcella[161]	Sp. nov	Valid	Robson & Young	Late Ordovician (middle Katian)	Red River Formation	Canada	A conulariid, a species of Conularia.
Gracilitheca astronauta[162]	Sp. nov	Valid	Valent & Fatka	Cambrian	Buchava Formation	Czech Republic	A member of Hyolitha (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Gracilitheca.
Gracilitheca mirabilis[163]	Sp. nov	Valid	Valent, Fatka & Marek	Cambrian	Buchava Formation	Czech Republic	A member of Hyolitha (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Gracilitheca.
Gracilitheca triangularis[163]	Sp. nov	Valid	Valent, Fatka & Marek	Cambrian	Buchava Formation	Czech Republic	A member of Hyolitha, a species of Gracilitheca.
Heteropistophyllum[159]	Gen. et comb. nov	Valid	Löser, Castro & Nieto	Early Cretaceous		Slovenia  Spain	A stony coral belonging to the group Heterocoeniina and the family Elasmocoeniidae. A new genus for "Pseudopistophyllum" quinqueseptatum Turnšek & Buser (1976).
Metaulastraea elae[159]	Sp. nov	Valid	Löser, Castro & Nieto	Early Cretaceous (Albian)		Spain	A stony coral, a species of Metaulastraea.
Microconchus utahensis[164]	Sp. nov	Valid	Zatoń, Taylor & Vinn	Early Triassic	Virgin Formation	United States	A member of Microconchida (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Microconchus.
Mitrodendron ultimus[159]	Sp. nov	Valid	Löser, Castro & Nieto	Early Cretaceous (Albian)		Spain	A stony coral, a species of Mitrodendron.
Nephrotheca betula[163]	Sp. nov	Valid	Valent, Fatka & Marek	Cambrian	Buchava Formation	Czech Republic	A member of Hyolitha, a species of Nephrotheca.
Nephrotheca sophia[162]	Sp. nov	Valid	Valent & Fatka	Cambrian	Buchava Formation	Czech Republic	A member of Hyolitha, a species of Nephrotheca.
Paraacanthogyra hispanica[159]	Sp. nov	Valid	Löser, Castro & Nieto	Early Cretaceous (Albian)		Spain	A stony coral, a species of Paraacanthogyra.
Paraconularia bachmanni[165]	Sp. nov	Valid	Barth et al.	Late Triassic		Germany	A conulariid, a species of Paraconularia.
Paranomalocaris[158]	Gen. et sp. nov	Valid	Wang, Huang & Hu	Cambrian	Wulongqing Formation	China	A member of Anomalocarididae (a group of animals with uncertain phylogenetic placement, possibly stem-arthropods). The type species is Paranomalocaris multisegmentalis.
Pleurodendron[159]	Gen. et sp. nov	Valid	Löser, Castro & Nieto	Early Cretaceous (Albian)		Spain	A stony coral belonging to the group Heterocoeniina and the family Elasmocoeniidae. The type species is Pleurodendron prebeticum.
Quasiaulacera[166]	Gen. et 2 sp. nov	Valid	Copper, Stock & Jin	Late Ordovician (Hirnantian)	Ellis Bay Formation	Canada	A stromatoporoid. Genus contains two species: Quasiaulacera stellata and Q. occidua.
Silingastraea valentina[159]	Sp. nov	Valid	Löser, Castro & Nieto	Early Cretaceous (Albian)		Spain	A stony coral, a species of Silingastraea.
Spinitheca[167]	Gen. et sp. nov	Valid	Novozhilova	Early Cambrian (Atdabanian)		Russia	A member of Hyolitha (a group of animals of uncertain phylogenetic placement, possibly molluscs). The type species is Spinitheca sysoievi.

References

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