Broiliellus

Broiliellus Temporal range: Early Permian
Restoration of Broiliellus olsoni
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Order:	†Temnospondyli
Family:	†Dissorophidae
Subfamily:	†Dissorophinae
Genus:	†Broiliellus Williston, 1914
Type species
†Broiliellus texensis Williston, 1914
Species
†B. arroyoensis DeMar, 1967 †B. brevis Carroll, 1964 †B. olsoni DeMar, 1967 †B. reiszi Holmes et al., 2013 †B. texensis Williston, 1914

Broiliellus is an extinct genus of dissorophoid temnospondyl within the family Dissorophidae. Broiliellus is most closely related to the genus Dissorophus, and both have been placed in the subfamily Dissorophinae. Broiliellus is known from three species from the Early Permian: the type species Broiliellus texensis and the species Broiliellus brevis and Broiliellus olsoni. A fourth species, Broiliellus novomexicanus, which was originally named Aspidosaurus novomexicanus, is now thought to fall outside the genus as a member of the subfamily Cacopinae.^[1]

Phylogeny

The phylogenetic analysis of Schoch (2012) found that Broiliellus was most closely related to Dissorophus. However, the three species of Broiliellus did not form their own clade, but rather a polytomy (unresolved evolutionary relationship) with Dissorophus.^[1]

Holmes, Berman & Anderson (2013) named a new species of Broiliellus, Broiliellus reiszi. They performed two phylogenetic analyses using the data matrix of Schoch (2012). In the second analysis the codings for "B." novomexicanus (i.e. Rio Arriba taxon) were not modified. This analysis yielded more resolved topology and a monophyletic Broiliellus with the exclusion of "B." novomexicanus. Below is the cladogram from that analysis:^[2]

Dissorophoidea

Micromelerpetontidae Micromelerpeton Amphibamidae Platyrhinops Doleserpeton Olsoniformes Trematopidae Fedexia Ecolsonia Tambachia Anconastes Phonerpeton Acheloma Dissorophidae Platyhystrix Aspidosaurus Brevidorsum Dissorophinae Dissorophus Broiliellus olsoni Broiliellus texensis Broiliellus brevis Broiliellus reiszi Cacopinae Conjunctio Rio Arriba taxon Admiral taxon Cacops aspidephorus Cacops morrisi Cacops woehri Kamacops Zygosaurus

References

1. Schoch, R. R. (2012). "Character distribution and phylogeny of the dissorophid temnospondyls". Fossil Record 15 (2): 121–137. doi:10.1002/mmng.201200010.
2. Holmes, R.; Berman, D. S.; Anderson, J. S. (2013). "A new dissorophid (Temnospondyli, Dissorophoidea) from the Early Permian of New Mexico (United States)". Comptes Rendus Palevol. doi:10.1016/j.crpv.2013.07.002.