Dipper

For other uses, see Dipper (disambiguation).
Dippers
American dipper
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Passeriformes
Suborder: Passeri
Family: Cinclidae
Sundevall, 1836
Genus: Cinclus
Borkhausen, 1797
Species

Dippers are members of the genus Cinclus in the bird family Cinclidae, named for their bobbing or dipping movements. They are unique among passerines for their ability to dive and swim underwater.

Description

White-throated dipper (C. cinclus)

Dippers are small, chunky, stout, short-tailed, short-winged, strong-legged birds. The different species are generally dark brown (sometimes nearly black), or brown and white in colour, apart from the rufous-throated dipper, which is brown with a reddish-brown throat patch. Sizes range from 14–22 cm in length and 40–90 g in weight, with males larger than females. Their short wings give them a distinctive whirring flight.[1][2][3] They have a characteristic bobbing motion when perched beside the water, giving them their name. While under water, they are covered by a thin, silvery film of air, due to small bubbles being trapped on the surface of the plumage.[4]

Distribution and habitat

Dippers are found in suitable freshwater habitats in the highlands of the Americas, Europe and Asia. In Africa they are only found in the Atlas Mountains of Morocco. They inhabit the banks of fast-moving upland rivers with cold, clear waters, though, outside the breeding season, they may visit lake shores and sea coasts.[2]

Adaptations

Unlike many water birds, dippers are generally similar in form to many terrestrial birds (for example they do not have webbed feet), but they do have some morphological and physiological adaptations to their aquatic habits. Their wings are relatively short but strongly muscled, enabling them to be used as flippers underwater. To reduce their buoyancy in water, the bones are solid instead of hollow.[5] They have dense plumage with a large preen gland for waterproofing their feathers. Relatively long legs and sharp claws enable them to hold on to rocks in swift water. Their eyes have well-developed focus muscles that can change the curvature of the lens to enhance underwater vision.[6] They have nasal flaps to prevent water entering their nostrils.

Their blood has a high haemoglobin concentration, allowing a greater capacity to store oxygen than terrestrial birds, and allowing them to remain underwater for up to at least thirty seconds,[2] whilst their basal metabolic rate is approximately one-third slower than typical terrestrial passerines of similar mass.[7] One small population wintering at a hot spring in Suntar-Khayata Mountains of Siberia feeds underwater when air temperatures drop below −55 °C (−67 °F).[8]

Behaviour

Food

Dippers forage for small animal prey in and along the margins of fast-flowing freshwater streams and rivers. They perch on rocks and feed at the edge of the water, but they often also grip the rocks firmly and walk down them beneath the water until partly or wholly submerged. They then search underwater for prey between and beneath stones and debris; they can also swim with their wings. The two South American species swim and dive less often than the three northern ones.[9] Their prey consists primarily of invertebrates such as the nymphs or larvae of mayflies, blackflies, stoneflies and caddisflies, as well as small fish and fish eggs. Molluscs and crustaceans are also consumed, especially in winter when insect larvae are less available.[2]

Breeding

Linear breeding territories are established by pairs of dippers along suitable rivers, and maintained against incursion by other dippers. Within their territory the pair must have a good nest site and roost sites, but the main factor affecting the length of the territory is the availability of sufficient food to feed themselves and their broods. Consequently, the length of a territory may vary from about 300 metres or 1,000 feet to over 2,500 metres or 8,200 feet.[2]

Dipper nests are usually large, round, domed structures made of moss, with an internal cup of grass and rootlets, and a side entrance hole. They are often built in confined spaces over, or close to, running water. The site may be on a ledge or bank, in a crevice or drainpipe, or beneath a bridge. Tree sites are rare.[2]

The usual clutch-size of the three northern dipper species is four or five; those of the South American species is not well known, though some evidence suggests that that of the rufous-throated dipper is two.[10] The incubation period of sixteen or seventeen days is followed by the hatching of altricial young which are brooded by the female alone for the next twelve to thirteen days. The nestlings are fed by both parents and the whole fledging period is about 20–24 days. Young dippers usually become independent of their parents within a couple of weeks of leaving the nest. Dippers may raise second broods if conditions allow.[2] Owing to the fact that bands inhibit their underwater movement, dippers are essentially unsuitable for ringing, so relatively little is known about maximum longevity or other aspects of their demographics, although records of a white-throated dipper living ten years and seven months do exist.[11]

Communication

Dippers’ calls are loud and high-pitched, being similar to calls made by other birds on fast rivers; the call frequencies lying within a narrow range of 4.0–6.5 kHz, well above the torrent noise frequency of <2 kHz.[12] Dippers also communicate visually by their characteristic dipping or bobbing movements, as well as by blinking rapidly to expose their pale upper eyelids as a series of white flashes in courtship and threat displays.[6][13]

Conservation

Dippers are completely dependent on fast-flowing rivers with clear water, accessible food and secure nest-sites. They may be threatened by anything that affects these needs such as water pollution, acidification and turbidity caused by erosion. River regulation through the creation of dams and reservoirs, as well as channelization, can degrade and destroy dipper habitat.[2]

Dippers are also sometimes hunted or otherwise persecuted by humans for various reasons. The Cyprus race of the white-throated dipper is extinct. In the Atlas Mountains dippers are claimed to have aphrodisiacal properties. In parts of Scotland and Germany, until the beginning of the 20th century, bounties were paid for killing dippers because of a misguided perception that they were detrimental to fish stocks through predation on the eggs and fry of salmonids.[2]

Despite threats to local populations, the conservation status of most dipper species is considered to be of least concern. The one exception, the rufous-throated dipper, is classified as vulnerable because of its small, fragmented and declining population which is threatened, especially in Argentina, by changes in river management.[14]

Taxonomy

Cinclus is the only genus in the family Cinclidae. The white-throated dipper and American dipper are also known in Britain and America, respectively, as the water ouzel (sometimes spelt "ousel") – ouzel originally meant the unrelated but superficially similar Eurasian blackbird (Old English osle). Ouzel also survives as the name of a relative of the blackbird, the ring ouzel.[15]

Species in taxonomic order

References

  1. ↑ Whistler, Hugh. (2007). Popular Handbook of Indian Birds. British Museum Natural History: London. 4th edition. ISBN 1-4067-4576-6
  2. 1 2 3 4 5 6 7 8 9 Tyler, Stephanie J.; & Ormerod, Stephen J. (1994). The Dippers. Poyser: London. ISBN 0-85661-093-3
  3. ↑ Robbins, C.S.; Bruun, B.; & Zim, H.S. (1966). Birds of North America. Western Publishing Company: New York.
  4. ↑ The Dippers
  5. ↑ White-throated Dipper - Tony Keene Birds
  6. 1 2 Goodge, W.R. (1960). "Adaptations for amphibious vision in the Dipper (Cinclus mexicanus)". J. Morphol. 107: 79–91. doi:10.1002/jmor.1051070106. PMID 13707012.
  7. ↑ Murrish, David E.; ‘Responses to Temperature in the Dipper, Cinclus mexicanus’; Comparative Biochemical Physiology; 1970, vol. 34, pp. 859 to 869
  8. ↑ Dinets, V. (2001) Brown Dipper Cinclus pallasi wintering in Suntar-Hayata Mountains of Yakutia. Ornitologia 29: 326 (in Russian with English summary).
  9. ↑ Tyler, S.J. (1994). "The Yungas of Argentina: in search of Rufous-throated Dippers Cinclus schulzi". Cotinga 2: 38–41.
  10. ↑ Salvador, S.; Narosky, S.; Fraga, R. (1986). "First description of the nest and eggs of the red-throated dipper in northwestern Argentina". Gerfaut 76: 63–66.
  11. ↑ Healy, Kevin, Guillerme, Thomas, Finlay, Sive, Kane, Adam, Kelly, Seán B. A., McClean, Deirdre, Kelly, David J., Donohue, Ian, Jackson, Andrew L. and Cooper, Natalie; ‘Ecology and mode-of-life explain lifespan variation in birds and mammals’; Proceedings of the Royal Society; June 2014(281); supplementary material
  12. ↑ J., Martens; Geduldig, G. (1990). "Acoustic adaptations of birds living close to Himalayan torrents". Proc. Int. 100 DO-G Meeting. Bonn: Current Topics Avian Biol. pp. 123–131.
  13. ↑ Creutz, G. (1966). Die Wasseramsel. A. Ziensen: Wittenburg Lutherstadt.
  14. ↑ BirdLife International (2007) Species factsheet: Cinclus schulzi. Downloaded from http://www.birdlife.org on 14/3/2008
  15. ↑ List of Old English bird names

External links

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