Dyakiidae

Dyakiidae
Five views of a shell of Dyakia salangana
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
(unranked):	clade Heterobranchia clade Euthyneura clade Panpulmonata clade Eupulmonata clade Stylommatophora informal group Sigmurethra clade limacoid clade
Superfamily:	Dyakioidea Gude & B. B. Woodward, 1921[1]
Family:	Dyakiidae Gude & B. B. Woodward, 1921[1]
Diversity
82 species (see text)
Synonyms
Sasakininae B. Rensch, 1930 Pseudoplectinae Thiele, 1934

Dyakiidae is a family of air-breathing land snails terrestrial pulmonate gastropod mollusks in the superfamily Dyakioidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).^[2]

Dyakiidae is the only family in the superfamily Dyakioidea.^[2] This family has no subfamilies.^[2]

Some of the species in this family are sinistral (left-handed) in their shell coiling.

Distribution

The family Dyakiidae is endemic to Sundaland in Southeast Asia.^[3]

Anatomy

In this family, the number of haploid chromosomes is between 26 and 30 (according to the values in this table).^[4]

The family also includes Quantula striata, the only known terrestrial gastropod to emit light.^[5]

The Digestive system characteristics are as follows. The buccal mass is small. The jaw is smooth. The stomach is very simple with weak muscles (as is the case in the majority of land snails).^[4] (These anatomical characteristics also include the family Staffordiidae which was considered part of the Dyakiidae at the time the study was done).

Genera

The family Dyakiidae includes the following 11 genera, with a total of 82 species (71 species until 2007^[6] + 11 new species in 2009^[7]):

• Asperitas Gray, 1857 - 14 species
• Bertia Ancey, 1887 - only one species: Bertia cambojiensis (Reeve, 1860)
• Dyakia Godwin-Austen, 1891 - type genus, 22 species
• Elaphroconcha Gude, 1911 - 10 species
• Everettia Godwin-Austen, 1891^[8] - 15 species
• Kalamantania Laidlaw, 1931 - only one species: Kalamantania whiteheadi (Godwin-Austen, 1891)^[9]
• Phuphania Tumpeesuwan, Naggs & Panha, 2007 - only one species: Phuphania globosa Tumpeesuwan, Naggs & Panha, 2007^[6]
• Pseudoplecta Laidlaw, 1932 - only one species: Pseudoplecta bijuga (Stoliczka, 1873)
• Quantula Baker, 1941 - only one species: Quantula striata (Gray, 1834)
• Rhinocochlis Thiele, 1931 - only one species: Rhinocochlis nasuta (Metcalfe, 1852)
• Sasakina Rensch, 1930 - four species

Cladogram

The following cladogram shows the phylogenic relationships of this family and superfamily with the other families within the limacoid clade:^[10]

limacoid clade	Staffordioidea  Staffordiidae  Dyakioidea  Dyakiidae  Gastrodontoidea  Pristilomatidae Chronidae Euconulidae Trochomorphidae Gastrodontidae Oxychilidae  Parmacelloidea  Trigonochlamydidae Parmacellidae Milacidae  Zonitoidea  Zonitidae  Helicarionoidea  Helicarionidae Ariophantidae Urocyclidae  Limacoidea  Vitrinidae Boettgerillidae Limacidae Agriolimacidae

References

1. Gude G. P. L. K. & Woodward B. B. (1921). "On Helicella, Férussac". Proceedings of the Malacological Society of London 14(5-6): 174-190. page 185.
2. Bouchet P., Rocroi J.-P., Frýda J., Hausdorf B., Ponder W., Valdés Á. & Warén A. (2005). "Classification and nomenclator of gastropod families". Malacologia: International Journal of Malacology (Hackenheim, Germany: ConchBooks) 47 (1-2): 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
3. Hausdorf, B. (1995). "A Preliminary Phylogenetic and Biogeographic Analysis of the Dyakiidae (Gastropoda: Stylommatophora) and a Biogeographic Analysis of Other Sundaland Taxa". Cladistics 11 (4): 359. doi:10.1111/j.1096-0031.1995.tb00095.x..
4. Barker G. M. (2001) Gastropods on Land: Phylogeny, Diversity and Adaptive Morphology. 1-146. In: Barker G. M. (ed.) (2001) The biology of terrestrial molluscs. CABI Publishing, Oxon, UK, cited pages: 139-144. ISBN 0-85199-318-4.
5. Isobe, M.; Uyakul, D.; Goto, T.; Counsilman, J. J. (1988). "Dyakia bioluminescence—1. Bioluminescence and fluorescence spectra of the land snail,D. Striata". Journal of Bioluminescence and Chemiluminescence 2 (2): 73–79. doi:10.1002/bio.1170020204. PMID 3213594..
6. Tumpeesuwan C., Naggs F. & Panha S. (31 August 2007) "A new genus and new species of dyakiid snail (Pulmonata: Dyakiidae) from the Phu Phan range, northeastern Thailand". Raffles Bulletin of Zoology 55(2): 363-369. PDF
7. Liew, T. S.; Schilthuizen, M.; Vermeulen, J. J. N. (2009). "Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo". Zoological Journal of the Linnean Society 157 (3): 515. doi:10.1111/j.1096-3642.2009.00526.x..
8. Godwin-Austen H. H. (1891) "On a collection of Land-shells made In Borneo by Mr. A. Everett, with Descriptions of supposed new species. Part II. Zonitidae and Helicidae". Proceedings of the Zoological Society of London for the year 1891: 22-47, p. 33.
9. Godwin-Austen H. H. (1891) "On a collection of Land-shells made In Borneo by Mr. A. Everett, with Descriptions of supposed new species. Part II. Zonitidae and Helicidae". Proceedings of the Zoological Society of London for the year 1891: 22-47, p. 24.
10. Hausdorf, B. (2000). "Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance events and long-distance dispersal". Journal of Biogeography 27 (2): 379. doi:10.1046/j.1365-2699.2000.00403.x. JSTOR 2656267..

Further reading

• Laidlaw F. F. (1931). "On a new sub-family Dyakiinae of the Zonitidae". Proceedings of the malacological Society of London 19: 190-201. abstract.
• Schileyko A. A. (2003). "Treatise on recent terrestrial pulmonate mollusks. 10. Ariophantidae, Ostracolethaidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae". Ruthenica, Supplement 2. 1309-1466.

External links

• "Dyakiidae". National Center for Biotechnology Information (NCBI).