Ennominae

Ennominae
Ourapteryx sp.
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Subphylum:	Hexapoda
Class:	Insecta
Order:	Lepidoptera
(unranked):	Macrolepidoptera
Superfamily:	Geometroidea
Family:	Geometridae
Subfamily:	Ennominae (Duponchel, 1845)
Tribes
Abraxini Angeronini Apeirini Apochimini Azelinini Baptini Bistonini (disputed) Boarmiini Bupalini (disputed) Caberini Campaeini (disputed) Cheimopteni Colotoini Cystidiini Desertobiini Ennomini Erannini (disputed) Gnophini (Duponchel, 1845) (disputed) Gonodontini (including Odontoperini) Lithinini (disputed) Macariini Melanolophiini (disputed) Nacophorini (disputed) Ourapterygini Phaseliini (disputed) Sphacelodini Theriini (disputed) Wilemanini and see text

Ennominae is the largest subfamily of the geometer moth family (Geometridae) with some 9,700 described species in 1,100 genera. They are usually fairly small moth species, though some (such as the Peppered Moth) grow considerably larger. This subfamily has a global distribution. It includes some species that are notorious defoliating pests.

The status of several tribes is debated. For example, the Boarmiini are sometimes massively expanded to include the Bistonini, Bupalini, Erannini, Gnophini, Melanolophini, Phaseliini and Theriini. The Nacophorini and perhaps the Campaeini might need to be merged with the Lithinini, and all three might warrant merging into the Ennomini. The group sometimes separated as Cassymini is tentatively included in the Abraxini here. The Alsophilinae, usually treated as a small subfamily in their own right, might simply be a specialized lineage of Boarmiini. ^[1]

Selected genera

Tribe Baptini

• Lomographa

Tribe Bupalini

• Bupalus

Tribe Caberini

• Cabera

Tribe Campaeini

• Campaea

Tribe Colotoini

• Colotois

Tribe Erannini

• Erannis

Tribe Gnophini

• Charissa
• Gnophos

Tribe Gonodontini

• Aethiopodes (sometimes in Odontopera)
• Odontopera

Genera incertae sedis

Numerous genera have hitherto not been definitely assigned to a tribe.^[2] These include:

Adalbertia Anavitrinelia Bichroma Chelotephrina Chemerina Chesiadodes Compsoptera Cyclomia Declana - Nacophorini? Enconista Eubarnesia Geolyces Hoplosauris - Ennomini? Hulstina Hypochrosis Liodesina Melanochroia Mericisca Metanema Metarranthis Miantochora Nepterotaea

Neritodes - Macariini? Onychora Ortaliella Orthocabera - Abraxini (Cassymini if distinct)? Orthofidonia - Boarmiini? Paraglaucina Parapheromia Prionomelia Probole Pseudocoremia - Boarmiini? Pterotaea Pyrinia Rhoptria Slossonia Sperrya Synglochis Toulgoetia Tracheops Tritocleis (extinct) Xenoecista Xylopteryx Zamarada - Abraxini (Cassymini if distinct)?

Footnotes

1. Holloway (1994), Young (2008)
2. See references in Savela (2008)

References

• Holloway, J.D. (1994): The Moths of Borneo: Family Geometridae, Subfamily Ennominae. London.
• Savela, Markku (2008): Markku Savela's Lepidoptera and some other life forms: Ennominae. Version of 2008-MAR-08. Retrieved 2008-JUL-21.
• Young, Catherine J. (2008): Characterisation of the Australian Nacophorini using adult morphology, and phylogeny of the Geometridae based on morphological characters. Zootaxa 1736: 1-141. PDF abstract and excerpt