Eurybacteria

Eurybacteria
Scientific classification
Domain:	Bacteria
Groups
Selenobacteria Fusobacteria Togobacteria

Eurybacteria is a taxon created by Cavalier-Smith,^[1] which includes several groups of Gram-negative bacteria. In this model, it is the ancestor of gram positive bacteria.^[2] Their endospores are characterized by producing and presenting external flagella or mobility by bacterial displacement.

Members

Specifically, it includes:

• Selenobacteria. For example, Sporomusa
  • Includes Heliobacteriales, for example, Heliobacterium
• Fusobacteria. For example, Leptotrichia and Fusobacterium
• Togobacteria (=Thermotogales). For example, Thermotoga.

In the standard classification, selenobacteria are usually included in the phylum firmicutes, whereas fusobacteria and togobacteria are classified as their own groups.

Relationships

The following graph shows Cavalier-Smith's version of the tree of life, indicating the status of eurybacteria.^[3]

[A]	Chlorobacteria  [B]  Hadobacteria  [C]   [D]  Cyanobacteria  [E]   [F]  Gracilicutes  [G]  Eurybacteria  [H] [I]  Endobacteria  [J]  Actinobacteria  [K] Neomura    [L]  Archaea  [M]  Eukarya

Legend: [A] Gram-negative with a peptidoglycan cell wall like Chlorosomee. [B] oxygenic Photosynthesis, Omp85 and four new catalases. [C] glycobacterial revolution: outer membrane with insertion of lipopolysaccharides, hopanoids, diaminopimelic acid, ToIC and TonB. [D] phycobilin chromophores. [E] Flagella. [F] Four sections: an amino acid in HSP60 and FtsZ and a domain in RNA polymerases β and σ. [G] Endospores. [H] Gram-Positive Bacteria: hypertrophy of the wall peptidoglycan, sortasa enzyme and a loss of the outer membrane. [I] glycerol 1-P dehydrogenase [J] proteasome and phosphatidylinositol. [K] Neomura revolution: Replacement of peptidoglycan by glycoproteins and lipoproteins. [L] reverse DNA gyrase and ether lipid isoprenoids. [M] Phagocytosis.

References

1. Cavalier-Smith T (2006). "Rooting the tree of life by transition analyses". Biol. Direct 1 (1): 19. doi:10.1186/1745-6150-1-19. PMC 1586193. PMID 16834776.
2. Cavalier-Smith T (January 2010). "Deep phylogeny, ancestral groups and the four ages of life". Philos. Trans. R. Soc. Lond., B, Biol. Sci. 365 (1537): 111–32. doi:10.1098/rstb.2009.0161. PMC 2842702. PMID 20008390.
3. Cavalier-Smith T (2006). "Cell evolution and Earth history: stasis and revolution" (PDF). Philos Trans R Soc Lond B Biol Sci 361 (1470): 969–1006. doi:10.1098/rstb.2006.1842. PMC 1578732. PMID 16754610.