List of national parks of Madagascar

This list of national parks of Madagascar includes all officially recognized protected areas as of 2015. The protected areas network of Madagascar is managed by the Madagascar National Parks Association (PNM-ANGAP). The network includes three types of protected areas: Strict Nature Reserves (IUCN category Ia), National Parks (IUCN category II) and Wildlife Reserves (IUCN category IV). At the 2003 IUCN World Parks Congress in Durban, the Malagasy President, Marc Ravalomanana, announced an initiative to more than triple the area under protection from approximately 4,200,791 acres (17,000.00 km²) to over 14,826,322 acres (60,000.00 km²) (from 3% to 10% of Madagascar's area). This "Durban Vision", as it has been dubbed, involved broadening the definition of protected areas in the country and legislation has been passed to allow the creation of four new categories of protected area: Natural Parks (IUCN category II), Natural Monuments (IUCN category III), Protected Landscapes (IUCN category V), and Natural Resource Reserves (IUCN category VI). As well as allowing these new objectives for protected areas management, the new legislation also provided for entities other than PNM-ANGAP to manage protected areas, such as government ministries, community associations, NGOs and other civil society organizations, and the private sector.

System of Protected Areas

The National Parks and Reserves of Madagascar (pre-2015)

The protection of natural sites in Madagascar was initiated under the French colonial authority in 1927. These original sites were reserved for scientific research and were not open to the public. In 1971, the Malagasy government undertook a project to protect 741,316 acres (3,000.00 km²) of mangrove forests, the first national effort to protect Madagascar's marine ecosystems. In 1986 the government of Madagascar, with support from the IUCN and the World Wildlife Fund, initiated a twelve-year process to review and assess existing protected areas and others requiring protection to create an initial list of Madagascar's conservation priority areas. The Association Nationale pour la Gestion des Aires Protégées (ANGAP), established in 1990, was the first government agency created with the express purpose of expanding and managing Madagascar's protected areas.^[1]

The creation of a national park system began in 1991 with the first major national policies for environmental protection and moved through three phases before concluding in 2002 with the establishment of the Systèmes des Aires Protégées de Madagascar (SAPM). As co-president of this commission, the World Wildlife Fund supports the government of Madagascar in managing the parks while also developing management partnerships with a broader variety of partners, including local communities, civil society and the private sector. In 2003, an additional 92 areas were identified as meriting the status of protected area; some of these have since been accorded an official protected status, while others are pending review. Ensuring the legal status and protection of the complete list of areas added to meet the Durban Vision commitment requires an updating of the law concerning protected areas, which was stalled following the political crisis of 2009.^[1]

On 17 September 2003 at the IUCN World Parks Congress in Durban, President Marc Ravalomanana announced an expansion of Madagascar's protected areas from 4,200,791 acres (17,000.00 km²) to over 14,826,322 acres (60,000.00 km²) (from 3% to 10% of Madagascar's area) over the next five years. The Malagasy government formed the Commission du Système des Aires Protégées de Madagascar (Commission for the Protected Areas System of Madagascar, SAPM) to work in partnership with the concerned government ministries (the Ministère de l’Environnement, des Eaux et Forêts [Ministry of Environment, Water and Forests] and the Ministère de l’Agriculture, de l’Elevage et de la Pêche [Ministry of Agriculture, Livestock and Fishery]). In March 2005, following a series of intensive collaborations with the UICN and other international and local experts, the government put in place the current system of classification and legal protection for Madagascar's protected areas.^[2]

World Heritage Sites

In 1999 the Tsingy de Bemaraha National Park was declared a UNESCO World Heritage Site.^[3] In 2007 six other national parks were voted in as a joint World Heritage Site under the name Rainforests of the Atsinanana. These six parks are Marojejy, Masoala, Ranomafana, Zahamena, Andohahela and Andringitra National Park.^[4] UNESCO placed the Rainforests of the Atsinanana on the list of World Heritage in Danger on 30 July 2010 following an increase in illegal logging in the parks since 2009 as a consequence of political crisis in the country.^[5]

Protected Areas

Strict Nature Reserves (Réserves Naturelles Intégrales)

Name	Photo	Location[6]	Date established[6]	Area[6]	Description[6][7][8]
Tsingy de Bemaraha Reserve		Melaky	1927	210,953 acres (853.7 km^2)	The area north of the Bemaraha National Park is reserved for scientific research and conservation of the highly endemic, diverse flora and fauna spanning the western coastal and inland transitional savanna ecosystems.
Betampona Reserve		Atsinanana	1927	7,215 acres (29.2 km^2)	This park is reserved for scientific research and conservation of the typical eastern lowland rainforest ecosystems. Many endemic plants with medicinal qualities are known to grow in this park, which shelters 93 bird species, of which 44 are endemic to Madagascar.
Tsaratanana Reserve		Sofia	1927	120,148 acres (486.2 km^2)	This park is reserved for scientific research and conservation of mountainous lowland and high altitude rainforest, including the highly endemic wildlife on the slopes of Mount Maromokotro, the highest in Madagascar.
Zahamena Reserve		Alaotra Mangoro, Analanjirofo, Atsinanana	1927	54,610 acres (221.0 km^2)	The central portion of Zahamena National Park is reserved for the conservation of several endemic and highly endangered plant and animal species. It is also the watershed of the Alaotra area, the highest producing agricultural area of the country.

National Parks (Parcs Nationaux)

Name	Photo	Location[6]	Date established[6]	Area[6]	Recreation Visitors (2013)[9]	Description[6][7][8]
Amber Mountain National Park		Diana	1958	44,973 acres (182.0 km^2)	10,770	This high altitude rain forest offers a cool, refreshing contrast to the hot plains surrounding the mountain. Countless waterfalls and streams cascade down the park's slopes and into its five lakes, providing drinking water for Antsiranana. The park is home to over 1,000 plant species and numerous lemur, amphibian and reptile species.
Andohahela National Park		Anosy	1939	187,849 acres (760.2 km^2)	156	Declared a World Heritage Site by UNESCO in 2007, this park includes typical southern spiny forest and the world's only rainforest south of the Tropic of Capricorn. Over 1,000 plant species grow in the park, including the triangle palm, which is nearly extinct in the wild.
Andringitra National Park		Haute Matsiatra	1927	76,998 acres (311.6 km^2)	3,156	Declared a World Heritage Site by UNESCO in 2007, this park offers spectacular vistas, strange rock formations, world class rock climbing and over 1,000 species of plants. The island's second tallest mountain, Imarivolanitra, lies within the park.
Ankarafantsika National Park		Boeny	2002	333,592 acres (1,350.0 km^2)	4,421	Containing dry deciduous forest, sandy savannas and numerous lakes and rivers, this park enjoys abundant animal diversity, particularly lemurs and birds.
Ankarana National Park		Diana	1956	45,035 acres (182.3 km^2)	12,643	This park contains limestone tsingy, dry deciduous forests, a sacred lake, and the largest network of caves and underground lakes and rivers anywhere in Africa.
Baie de Baly National Park		Boeny	1997	141,200 acres (571.4 km^2)	100	This park protects the unique ecosystem at the intersection of the marine reefs, the mangrove-lined coastal habitats and the dry deciduous forests of the northwest, providing refuge for species such as the dugong and the big-headed turtle. Over 86% of all waterfowl species in Madagascar are found in this park, where many of them have established colonies.
Isalo National Park		Ihorombe	1962	201,490 acres (815.4 km^2)	28,375	The most visited park in Madagascar, Isalo preserves a sandstone massif that has eroded over time to form dramatic and colorful canyons, plateaus and valleys. Seventeen rivers cut through the arid landscape, which is dotted with palms and inhabited by ringtail lemurs.
Kirindy Mitea National Park		Menabe	1997	178,410 acres (722.0 km^2)	unknown	This park protects the unique wildlife of the hot and dry transitional landscape between the western and southern ecosystems. Although diversity of species is lower relative to the eastern rainforests, endemism is very high, constituting 100% of its reptiles and amphibians, 91% of mammals in the park, 70%, of bird species and 70% of the park's plant species.
Lokobe National Park		Diana	1913	3,763 acres (15.2 km^2)	n/a	Extending over a portion of the island of Nosy Be and encompassing forest, coastal and marine ecosystems, this park protects one of the last examples of old growth coastal forest particular to the Sambirano area. Its status was changed from Strict Nature Reserve to National Park in June 2014.[10]
Mananara Nord National Park		Analanjirofo	1989	355,832 acres (1,440.0 km^2)	100	Madagascar's first marine park comprises the marine ecosystems around a cluster of three islands. It was the first protected area to be included in the UNESCO global network of biosphere reserves. The coral reefs here are among the best in Madagascar.
Andasibe-Mantadia National Park		Alaotra-Mangoro	1989	38,252 acres (154.8 km^2)	25,684	Connected by a forest corridor to Zahamena National Park, Mantadia shelters the island's largest lemur, the critically endangered indri. A wide variety of rare orchids bloom in this high altitude rainforest.
Marojejy National Park		Sava	1952	148,387 acres (600.5 km^2)	1,362	Declared a World Heritage Site by UNESCO in 2007, Marojejy National Park protects the primary lowland coastal rainforest on and surrounding the sacred mountain of Marojejy. Numerous endemic bird species and 33% of all amphibian species found in Madagascar inhabit the park.
Masoala National Park, including Nosy Mangabe Reserve		Sava, Analanjirofo	1997	594,338 acres (2,405.2 km^2)	2,480	Declared a World Heritage Site by UNESCO in 2007, Masoala contains four parcels of primary rainforest ranging from sea level to 1,300 meters, and three adjacent marine parcels. Over 50% of all the plant and animal species found in Madagascar inhabit this park, which is the sole habitat of the endangered red ruffed lemur.
Midongy du sud National Park		Atsimo-Atsinanana	1997	474,932 acres (1,922.0 km^2)	unknown	Containing an especially high number of endemic medicinal plants, this mid to high altitude rainforest park also protects 14 endangered endemic animal species and groves of precious ebony and pallisander.
Ranomafana National Park		Haute Matsiatra, Vatovavy-Fitovinany	1991	102,798 acres (416.0 km^2)	21,032	Declared a World Heritage Site by UNESCO in 2007, this park protects transitional rainforest between typical eastern and southern ecosystems. It is home to 115 bird species (30 unique to the park) and the golden bamboo lemur. The hot springs here have long been visited for their healing properties.
Sahamalaza National Park		(marine) - Sofia	2007	64,247 acres (260.0 km^2)	unknown	Sahamalaza, also called Iles Radama, was initially a UNESCO Biosphere Reserve before being named a national park in 2007. About half of the park is underwater and comprises a network of coral reefs, while the remainder includes mangroves along the northwest coast and 75 square kilometers of some of the last remaining dry littoral forest on the west coast. The critically endangered Sahamalaza sportive lemur (fewer than 100 remaining) and Sclater's black lemur (the world's only blue-eyed non-human primate) are endemic to this forest.
Tsimanampetsotse National Park		Atsimo-Andrefana	1927	106,750 acres (432.0 km^2)	1,186	This park protects Lake Tsimanampetsotsa, Madagascar's only saline lake. Between 75 and 90% of the flora and fauna in the park are endemic. The park is home to a blind cave-dwelling goby and a large colony of flamingos year-round.
Tsingy de Bemaraha National Park		Melaky	1997	178,756 acres (723.4 km^2)	9,561	This park was declared a UNESCO World Heritage Site in 1990. It contains the largest of Madagascar's limestone tsingy fields, as well as dry forest, bush, rainforest and savanna. The biodiversity contained in this park is among the richest of any protected natural area in the world: 87% of the plants and animals are endemic to Madagascar, and 45% are uniquely endemic to the region around the park.
Tsingy de Namoroka National Park		Boeny	2002	54,924 acres (222.3 km^2)	unknown	The dramatic landscapes preserved in this park include tsingy, canyons, caves and pools. Connected to the Bay de Baly National Park, Namoroka's status was changed from a Strict Nature Reserve to a National Park in 2002.
Zahamena National Park		Alaotra Mangoro, Analanjirofo, Atsinanana	1997	104,526 acres (423.0 km^2)	unknown	Declared a World Heritage Site by UNESCO in 2007, this eastern rainforest park is home to a rich diversity of endemic animal species, including 112 species of birds, 29 fish species, 62 different amphibian, and 46 reptiles. On the 48 species of mammals counted, 13 are lemurs.
Zombitse-Vohibasia National Park		Atsimo-Andrefana	1997	89,719 acres (363.1 km^2)	2,822	The dry, hot southwestern forests protected by this park are intercut with rivers that support a variety of endemic wildlife, including 47% of all endemic bird species in Madagascar.

Wildlife Reserves (Réserves Spéciales)

Name	Photo	Location[6]	Date established[6]	Area[6]	Description[6][7][8]
Ambatovaky Reserve		Analanjirofo	1958	148,387 acres (600.5 km^2)	Ambatovaky is the largest protected lowland rainforest after Masoala and the largest of Madagascar's special reserves. 75% of the 291 plant species here are endemic. The reserve harbors 11 species of lemurs, 110 bird species, 113 species of amphibian and 34 fish species.
Amber Forest Reserve		Diana	1958	11,886 acres (48.1 km^2)	The reserve adjoins the Amber Mountain National Park and serves to protect the wealth of biodiversity and endemism common to both areas, including 1,020 plant species, of which many have medicinal properties.
Ambohijanahary Reserve		Menabe, Melaky	1958	61,159 acres (247.5 km^2)	The park protects the highly threatened sclerophyllous forest in the transitional zone between the western and cental ecosystems. Altitudes vary from 200 to 900 meters above sea level, with dramatic variation in plant life following the increase in elevation.
Ambohitantely Reserve		Analamanga	1982	13,838 acres (56.0 km^2)	Ambohitantely protects the last remaining fragment of the forest that formerly covered the central highlands of Madagascar. This forest contains a blend of high altitude rainforest evergreen species interspersed with vegetation found in the mid-altitude sclerophyll forests to the west, including the threatened Manambe palm.
Analamazoatra Reserve (Périnet)		Alaotra Mangoro	1989	38,252 acres (154.8 km^2)	This reserve is connected to Mantadia National Park, forming the larger Andasibe-Mantadia protected area that shelters aye-aye, over 100 species of orchids, numerous ferns and palms, and the indri.
Analamerana Reserve		Diana, Sava	1956	85,746 acres (347.0 km^2)	This is the only reserve protecting the habitat of the critically endangered Perrier's sifaka. The reserve also shelters the eight rarest bird species in the world in its tsingy and forested canyons, where three of the island's seven baobab species grow.
Andranomena Reserve		Menabe		15,864 acres (64.2 km^2)	Protecting the typical ecosystem of the Menabe region, this reserve's plant life is 80% endemic and includes the Grandidier's baobab and two other baobab species growing in their natural forest habitat. The Verreaux's sifaka and four other endangered lemur species inhabit the park.
Anjanaharibe-Sud Reserve		Sava	1958	79,296 acres (320.9 km^2)	This rainforest reserve ranges in altitude from 500 to 2,000 meters. It shelters immense plant and animal diversity, much of which is endemic, including 125 species of birds, 12 lemur species, 53 species of amphibians, 40 reptile species, 30 species of palms, and 300 fern species, of which four are found only within the reserve. This is the last habitat of Takhtajania, an angiosperm plant that evolved to its present state 120 million years ago.
Bemarivo Reserve		Melaky	1956	28,590 acres (115.7 km^2)	The rivers, lakes and swamps of Bemarivo shelter 15 species of mammals, 20 species of reptile, six species of lemurs and three species of carnivores including the endemic fossa. Almost 47% of the reserve's 73 bird species are endemic to Madagascar, while another 30% are endemic to the Indian Ocean region.
Beza Mahafaly Reserve		Atsimo-Andrefana	1978	1,483 acres (6.0 km^2)	Beza Mahafaly contains the only protected gallery forest in Madagascar, which runs along the Sakamena River. A large portion of southwest Madagascar's radiated tortoise population lives in the park, alongside ring-tailed lemurs, sifaka, mouse lemurs and sportive lemurs.
Bora Reserve		Sofia	1956	11,962 acres (48.4 km^2)	The Bora forest represents the transitional ecosystem between the dry western forests and the eastern rain forests at the northern end of the island. Numerous bird species typically inhabiting either type of forest can be found here, of which 20 are endemic.
Cap Sainte Marie Reserve		Androy	1962	4,324 acres (17.5 km^2)	Also known as Cape Vohimena, this reserve protects the highly endangered radiated tortoise and the unique dwarf vegetation of this arid, desert-like region. Egg shell fragments of the extinct elephant bird litter the sand. Humpback whales migrate along the cape shore from August to November. The park's turtle population averages 3,000 turtles per square kilometer, one of the highest densities in the world.
Kalambatritra Reserve		Ihorombe, Anosy	1959	69,819 acres (282.5 km^2)	Separated from the Midongy du Sud National Park by a 20 kilometer savanna corridor, the Kalambaritra Reserve is unique in encompassing the ecosystems of the eastern lowland rainforest, southern spiny forest and central temperate forests in a single park, as well as expanses of marshlands that protect numerous fish species. It is rich in bird, lemur and amphibian species; threatened lemurs include the collared brown lemur and the common brown lemur.
Kasijy Reserve		Betsiboka	1956	48,927 acres (198.0 km^2)	This reserve is distinguished by the diversity of the ecosystems protected on this flat, hot plain, including swamps, savannas, steppes, dry deciduous forests and dense temperate forests. Among the 67 species of birds found here, 43% are endemic to Madagascar and 34% are endemic to Madagascar, Comoros and the Mascarene Islands.
Mangerivola Reserve		Atsinanana	1958	32,136 acres (130.0 km^2)	The low, medium and high altitude rainforests protected within the Mangerivola Reserve are home to 100 bird species, of which 63 are endemic to Madagascar and 23 are endemic to the region. These endemic species include the Madagascan serpent eagle - one of the rarest birds in the world - and the one of the rarest owls, the endemic red owl. Half the chameleons found in the park are considered rare.
Maningoza Reserve		Melaky	1956	19,521 acres (79.0 km^2)	The dense dry forest of Maningoza is one of the best examples of this type of ecosystem, with 78% of its plant species endemic to Madagascar. It is one of the last examples of tropical dry forest on ironitic soil that formerly covered much of western Madagascar.
Manombo Reserve		Atsimo-Atsinanana	1962	13,146 acres (53.2 km^2)	The lowland rainforest of Manombo is home to a wide variety of animal species, of which 90% are endemic. It is the only protected area providing shelter to the endangered poisonous Bernhard's mantella frog, and hosts 52 species of endemic snails, the greatest diversity of any protected area in Madagascar. The reserve's flora include one of two species of the tree genus Allantospermum, the other being in Borneo; it is also the only protected habitat of the endemic Hintsia bijuga tree, formerly more widespread in the eastern rainforest.
Manongarivo Reserve		Diana	1956	80,890 acres (327.4 km^2)	The reserve protects a forest typical of the Sambirano area that shelters 11 plant species endemic to the reserve, as well as four types of lemur, 103 species of birds, 31 amphibian species, and 39 reptile species. The landscape of the reserve is dramatic. It ranges from 150 to 1,200 meters in altitude, encompassing low and high altitude dry deciduous forest growing over a stony massif of gneiss, granite and basalt that juts up abruptly to form mountains and cliffs.
Marotandrano Reserve		Sofia, Alaotra-Mangoro	1956	104,278 acres (422.0 km^2)	The deep, narrow valleys of Marotandrano are covered in dense mid-altitude rainforest and shelter 140 species of birds (56 endemic), 19 species of amphibian, 16 reptile species and 12 species of lemurs, including the black-and-white ruffed lemur.
Pic d'Ivohibe Reserve		Ihorombe	1964	8,532 acres (34.5 km^2)	Connected to Andringitra National Park, Ivohibe reserve shelters 77 species of birds in its dense low and mid altitude rain forests. The rivers running down its peak and surrounding slopes provide irrigation to the farms of surrounding communities.
Tampoketsa Analamaitso Reserve		Sofia	1958	42,379 acres (171.5 km^2)	One of the last examples of transitional forest between western and eastern forest ecosystems, Tampoketsa includes mid-altitude rainforest and dense dry deciduous forests. Almost half of the reserve's fauna is endemic, including 23 out of the park's 24 reptile species; the reserve also has unusually diverse snail and beetle populations.

All other Protected Areas

Name	Photo	Location[11]	Date established[11]	Area[11][12]	Description
Fandriana Marolambo Forest Corridor		Atsinanana, Vakinankaratra, Amoron'i Mania	2015[13]	480,098 acres (1,942.9 km^2)	This protected area, which comprises savannas and secondary forest interspersed with degraded forest and farms, forms a corridor between several of the UNESCO World Heritage Ala Atsinanana parks. Among the plants found in this area, 26% of the families and 95% of the species are endemic; 13 lemur species and many endemic amphibian and reptile species are also found here.[14]
Mahavavy-Kinkony Protected Area		Boeny	2006	682,010 acres (2,760.0 km^2)	Mahavavy-Kinkony encompasses dense dry forest, freshwater lake, saline bay and riverine delta ecosystems that are home to several critically endangered endemic species.[15]
Montagne des Français Protected Area		Diana	2013[16]	15,053 acres (60.9 km^2)[16]	This protected area to the east of Antsiranana is a dense and dry semi-deciduous forest covering a limestone plateau with canyons. A unique baobab species is endemic to this area, as are three vascular plant families and several animal species.[16]
Nosy Hara Marine and Coastal Protected Area		Diana	2012	452,477 acres (1,831.1 km^2)	This island is the sole habitat of the world's smallest chameleon. The forested island features patches of tsingy and is ringed by very well preserved coral reefs.
Nosy Tanikely National Marine Park		Diana	2011[17]	773 acres (3.1 km^2)	This marine park protects a network of coral reefs that host a variety of endemic and endangered Indian Ocean aquatic species, including sea turtles.[17] There were 22,051 visitors to Nosy Tanikely in 2013.
Nosy Ve Marine and Coastal Protected Area		Atsimo-Andrefana	2015[13]	4,853 acres (19.6 km^2)	This park protects a network of coral reefs containing approximately 140 species of coral and over 240 species of fish, including fish of the very rare Coelacanth family. Dolphins, sea turtles and humpback whales are also common here.[18]

See also

• List of World Heritage Sites in Madagascar

References

1. "WWF: 50 ans à Madagascar" (PDF) (in French). World Wildlife Fund. 2013. Retrieved 4 June 2015.
2. "Système des Aires Protégées de Madagascar" (in French). Ministre de l’Environnement, de l’Ecologie, de la Mer et des Forêts. 2015. Retrieved 4 June 2015.
3. "Report of the World Heritage Committee, 14th Session". UNESCO. Retrieved 1 June 2011.
4. "World Heritage Convention: Madagascar". United Nations Educational and Cultural Organization (UNESCO). Retrieved 25 April 2011.
5. "World Heritage Committee inscribes Rainforests of Atsinanana (Madagascar) on List of World Heritage in Danger". UNESCO. Retrieved 25 April 2011.
6. Madagasikara National Parks (2014). "Madagascar National Parks". Parcs-madagascar.com. Archived from the original on March 1, 2014. Retrieved 13 September 2014.
7. "Madagascar Travel Guide". Travelmadagascar.org. 2014. Retrieved 13 September 2014.
8. Butler, Rhett (2012). "Parks in Madagascar". Wildmadagascar.org. Retrieved 13 September 2014.
9. Office National du Tourisme de Madagascar (May 2014). "Le Site du Magazine Officiel du tourisme a Madagascar" (in French). Travelmadagascar.org. Archived from the original on May 23, 2014. Retrieved 13 September 2014.
10. "Nosy-Be: Le Parc National Lokobe est ouvert à l'écotourisme à partir du 28 juin 2014" (in French). Sobika.com. 24 June 2014. Retrieved 23 September 2014.
11. "The new protected areas (NAP)". Madagascar Biodiversity Fund. 2009. Retrieved 3 June 2015.
12. "AP MNP et Extensions" (in French). Ministre de l’Environnement, de l’Ecologie, de la Mer et des Forêts. 2015. Retrieved 3 June 2015.
13. "Conseil du Gouvernement du Mardi 21 Avril 2015 au Palais d’Etat de Mahazoarivo" (in French). Sobika. 22 April 2015. Retrieved 4 June 2015.
14. Roelens, Jean-Baptiste; Vallauri, Daniel; Razafimahatratra, Appolinaire; Rambeloarisoa, Gérard; Razafy, Fara Lala (2010). "Restauration des Paysages Forestieres: Cinq ans de réalisations à Fandriana-Marolambo (Madagascar)" (PDF). World Wildlife Fund. Retrieved 4 June 2015.
15. "Mahavavy-Kinkony" (in French). Ministere de l’Environnement, de l’Ecologie, de la Mer et des Forêts. 2015. Retrieved 3 June 2015.
16. "Montagne des Français". Madagascar Biodiversity Fund. 2014. Retrieved 13 September 2014.
17. "Le Parc National Marin de Nosy Tanikely" (in French). La Tribune de Diego. 17 July 2012. Retrieved 4 June 2015.
18. "Information scientifique (écologique et biologique marines) pour l’APM Nosy Ve Androka" (PDF) (in French). CBD International. 2015. Retrieved 4 June 2015.