Urtinotherium

Urtinotherium Temporal range: Late Eocene to Early Oligocene, 37–30 Ma PreЄ Є O S D C P T J K Pg N
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Synapsida
Class:	Mammalia
Order:	Perissodactyla
Family:	†Hyracodontidae
Subfamily:	†Indricotheriinae
Genus:	†Urtinotherium Chow & Chiu, 1963
Species
U. incisivum Chow & Chiu, 1963 U. yagouense (Qui et al., 2004) U. parvum (Chow, 1958)[1]
Synonyms
Turpanotherium Qui & Wang, 2007 Species synonyms Urtinotherium yagouense: Paraceratherium yagouense Qui et al., 2004 Turpanotherium yagouense (Qui et al., 2004) Turpanotherium elegans Qui & Wang, 2007 Urtinotherium parvum: Indricotherium parvum Chow, 1958 Indricotherium qujigensis Tang, 1978

Urtinotherium (meaning "Urtyn beast") is an extinct genus of indricothere hyracodontid mammals. It was a large animal that was closely related to Paraceratherium, and found in rocks dating from the Late Eocene period. The remains were first discovered in the Urtyn Obo region in Inner Mongolia, which the name Urtinotherium is based upon. Other referred specimens are from northern China.^[2]

Description

Urtinotherium was a representative of the large subfamily Indricotheriinae, and almost reached the proportions of Paraceratherium. It is known from several findings in central and eastern Asia, though no complete skeletons are known. The holotype fossil (catalog number IVPP V.2769) includes a complete lower jaw 71.5 cm (28.1 in) in length, comparable to the smallest known of Paraceratherium. The jaw is wedge shaped, very low in height, and with elongated branches. The symphysis was solid and extended to the beginning of the second premolars. The jaws dentition is complete, having three front incisors and canine. The first two incisors protrude forward with a crown length of 4.9 cm (1.9 in), with forms similar to daggers. The other incisors and canines had much smaller crowns. Between each tooth is a small space, similar to its phylogenetically primitive relatives such as Juxia. Its back teeth, which are separated from the front teeth by a small diastema, consist of four premolars and three molars. These are similar in structure to those of Paraceratherium, with small premolars and larger molars. The latter have low coronas (are brachyodont) and had few enamel folds.^[3]

Distribution

The remains of Urtinotherium are found mainly in the eastern and central Asia, consisting mostly of jaw fragments and isolated teeth. The holotype jaw was found in the early 1960s in the Urtyn-Obo Formation of Inner Mongolia, which dates from the Late Eocene.^[3] Additional findings came from Yunnan province in China and Late Eocene deposits of Khoer-Dzam in Mongolia. Further findings have been made in Saissansee Aksyir Svita basin in eastern Kazakhstan, also of the same age.^[2] Urtinotherium likely survived into the Early Oligocene, as fossils probably from it have been found in the Mera Formation of Transylvania, Romania.^[4]

Classification

Urtinotherium pertains to the Hyracodontidae subfamily Indricotheriinae. These in turn are part of the superfamily Rhinocerotoidea and therefore represent close relatives of modern rhinoceroses. The hyracodontids are distinguished by the formation of large sharp incisors in their upper and lower jaws, while rhinoceroses only have a two on the lower jaw. Urtinotherium was thought of by Leonard Radinsky to be a transitional form between earlier indricotheres, like Juxia, and later forms, such as Paraceratherium and Indricotherium (now Paraceratherium transouralicum).^[5]

This genus represents a primitive form of indricothere that developed in the late Eocene. It is possibly descended from Juxia, which lived during the Middle Eocene in northern China, and which shares with it having a full set of teeth in its jaws. However, Urtinotherium differs by its larger body size and greater specialization in the incisors.^[6] Urtinotherium and later Paraceratherium form the most derived section of the largest known rhinocerotoid lineage. In their specialization emphasizes that compared to Urtinotherium, they possessed significantly reduced dentition with only a couple of incisors in the lower jaw.^[7]

The first description of Urtinotherium was published in 1963 by Zhou Ming-Zhen and Chiu Chan-Siang, based on the lower jaw. The type species is recognized as Urtinotherium incisivum. The genus name comes from the location, and from the Ancient Greek word θηρίον (therion) meaning "beast." The species name refers to its elongated incisivos.^[3]

References

1. Wang, H.-B.; Bai, B.; Gao, F.; Huang, W.-C.; Wang, Y.-K. (2013). "New eggysodontid (Mammalia, Perissodactyla) material from the Paleogene of the Guangnan Basin, Yunnan Province, China" (PDF). Vertebrata PalAsiatica 51 (4): 307–322.
2. Bayshashanov, U.B.; Lucas, S.G. (2001). "The giant rhinoceros Urtinotherium from the upper Eocene of the Zayan Basin, Kazakhstan". Zoological journal of Kazakhstan: 185–187. ISSN 1024-7688.
3. Chow, M.; Chiu, C.-S. (1963). "New genus of giant rhinoceros from Oligocene of Inner Mongolia" (PDF). Vertebrata PalAsiatica 7 (9): 230–239.
4. Codrea, V.A. (2000). Rinoceri și Tapiri Terțiari din România [Tertiary Rhinoceroses and Tapirs of Romania] (in French). Presa Universitara Clujeana. pp. 145–147. ISBN 973-8095-18-2.
5. Radinsky, L.B. (1966). "The families of the Rhinocerotoidea (Mammalia, Perissodactyla)". Journal of Mammalogy 47 (4): 631–639. doi:10.2307/1377893. JSTOR 1377893.
6. Chow, M.; Chiu, C.-S. (1964). "An eocene giant rhinoceros". Vertebrata PalAsiatica (8): 264–268.
7. Lucas, S.G.; Sobus, J.C. (1989). "The Systematics of Indricotheres". In Prothero, David R.; Schoch, Robert M. The Evolution of Perissodactyls. Oxford University Press. pp. 358–378. ISBN 978-0-19-506039-3. OCLC 19268080.