Upeneus

Upeneus
Upeneus parvus
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	Perciformes
Family:	Mullidae
Genus:	Upeneus G. Cuvier, 1829[1]
Type species
Mullus vittatus Forsskål, 1775
Synonyms
Hypeneus Agassiz, 1846 Upeneoides Bleeker, 1849 Megalepis Bianconi, 1857 Pennon Whitley, 1941

Upeneus is a genus of goatfishes native to the Atlantic, Indian and Pacific oceans.

Species

There are currently 37 recognized species in this genus:

Species[2]	Taxon authority	Common name	Environment & range[3]
Upeneus asymmetricus	Lachner, 1954	Asymmetrical goatfish	Philippines, Indonesia
Upeneus australiae	B. J. Kim & Nakaya, 2002	Australian goatfish	Eastern Indian Ocean and Western Pacific; Australia
Upeneus davidaromi	Golani, 2001	Darom's goatfish	Red Sea; deep-water (150–500 m)
Upeneus doriae	Günther, 1869	Gilded goatfish	Persian Gulf and the Gulf of Oman
Upeneus filifer	J. D. Ogilby, 1910	Pennant goatfish	West-central Pacific
Upeneus francisi	J. E. Randall & Guézé, 1992	Francis' goatfish	Norfolk Island and New Zealand
Upeneus guttatus	F. Day, 1868	Two-tone goatfish	Indo-West Pacific: the Red Sea to South Africa and to northern Australia
Upeneus heemstra	Uiblein & Gouws, 2014 [4]	Heemstra goatfish	Western Indian Ocean: Coast of Africa to India & Sri Lanka
Upeneus indicus	Uiblein & Heemstra, 2010	Tall-fin goatfish	Western India
Upeneus itoui	M. Yamashita, Golani & Motomura, 2011 [5]	Oriental goatfish	Southern Japan
Upeneus japonicus	Houttuyn, 1782	Japanese goatfish	Japan north to Korea and south to Taiwan and Hong Kong
Upeneus lombok	Uiblein & W. T. White, 2015 [6]	Lombok goatfish	Lombok, Indonesia
Upeneus luzonius	D. S. Jordan & Seale, 1907	Dark-barred goatfish	Pacific waters around the Philippines, Indonesia and northwestern Australia
Upeneus margarethae	Uiblein & Heemstra, 2010	Margaretha's goatfish	Indian Ocean
Upeneus mascareinsis	Fourmanoir & Guézé, 1967	Mascarene goatfish	Madagascar, Mozambique, Réunion, Western Indonesia
Upeneus moluccensis	Bleeker, 1855	Goldband goatfish	Indo-West Pacific
Upeneus mouthami	J. E. Randall & Kulbicki, 2006	Moutham's goatfish	Western Central Pacific
Upeneus niebuhri	Guézé, 1967 [4]	Niebuhr’s goatfish	Gulf of Suez
Upeneus nigromarginatus	A. R. Bos, 2014 [7]	Black-margined goatfish	Philippines (Davao Gulf)
Upeneus oligospilus	Lachner, 1954	Short-fin goatfish	Persian Gulf
Upeneus parvus	Poey, 1852	Dwarf goatfish	Western Atlantic
Upeneus pori	Ben-Tuvia & Golani, 1989	Por's goatfish	Red Sea to southern Oman
Upeneus quadrilineatus	C. T. Cheng & C. X. Wang, 1963	Four-stripe goatfish	Southern Japan to southern China
Upeneus randalli	Uiblein & Heemstra, 2011		Persian Gulf
Upeneus saiab	Uiblein & Lisher, 2013 [8]	Saiab goatfish	Indian Ocean off coast of Mozambique
Upeneus seychellensis	Uiblein & Heemstra, 2011	Tailstripe goatfish	Seychelles Bank in the western Indian Ocean
Upeneus stenopsis	Uiblein & McGrouther, 2012	Narrowtail goatfish	Western Pacific Ocean
Upeneus suahelicus	Uiblein & Heemstra, 2010	Swahili goatfish	Kenya to South Africa, Eritrea
Upeneus subvittatus	Temminck & Schlegel, 1843	Deep-water goatfish	Japan to Indonesia
Upeneus sulphureus	G. Cuvier, 1829	Sulphur goatfish	Indo-West Pacific
Upeneus sundaicus	Bleeker, 1855	Ochrebanded goatfish	Pakistan, India and Sri Lanka
Upeneus supravittatus	Uiblein & Heemstra, 2010	Long-fin goatfish	Sri Lanka, southern India
Upeneus taeniopterus	G. Cuvier, 1829	Finstripe goatfish	Indo-Pacific
Upeneus torres	Uiblein & Gledhill, 2015 [9]	Torres goatfish	Australia, Vanuatu
Upeneus tragula	J. Richardson, 1846	Freckled goatfish	Indo-West Pacific
Upeneus vanuatu	Uiblein & Causse, 2013 [10]	Vanuatu goatfish	Pacific Ocean, only known from Vanuatu
Upeneus vittatus	Forsskål, 1775	Yellowstriped goatfish	Indo-Pacific
Upeneus xanthogrammus	C. H. Gilbert, 1892		Eastern Pacific

References

1. Cuvier, G. (1829): Le Règne Animal, distribué d'après son organisation, pour servir de base à l'histoire naturelle des animaux et d'introduction à l'anatomie comparée. Edition 2. 2: i-xv + 1-406.
2. Uiblein, F. & Heemstra, P.C. (2010): A taxonomic review of the western Indian Ocean goatfishes of the genus Upeneus (Family Mullidae), with descriptions of four new species. Smithiana, Publications in Aquatic Biodiversity, Bulletin, 11: 35-71.
3. Lieske, E. & Myers, R. (1994): Collins Pocket Guide. Coral reef fishes. Indo-Pacific & Caribbean including the Red Sea. Haper Collins Publishers, 400 p.
4. Uiblein, F. & Gouws, G. (2014): A new goatfish species of the genus Upeneus (Mullidae) based on molecular and morphological screening and subsequent taxonomic analysis. Marine Biology Research, 10 (7): 655-681.
5. Yamashita, M., Golani, D. & Motomura, H. (2011). "A new species of Upeneus (Perciformes: Mullidae) from southern Japan". Zootaxa 3107: 47–58.
6. Uiblein, F. & White, W.T. (2015). "A new goatfish of the genus Upeneus (Mullidae) from Lombok, Indonesia and first verified record of U. asymmetricus for the Indian Ocean" (PDF). Zootaxa 3980 (1): 51–66. doi:10.11646/zootaxa.3980.1.3.
7. Bos, A.R. (2014). "Upeneus nigromarginatus, a new species of goatfish (Perciformes: Mullidae) from the Philippines" (PDF). Raffles Bulletin of Zoology 62: 745–753.
8. Uiblein, F. & Lisher, M. (2013): A new goatfish of the genus Upeneus (Mullidae) from Angoche, northern Mozambique. Zootaxa, 3717 (1): 85–95.
9. Uiblein F. & Gledhill D.C. (2015). "A new goatfish of the genus Upeneus (Mullidae) from Australia and Vanuatu, with inter- and intraspecific comparisons". Marine Biology Research 11 (5): 475–491. doi:10.1080/17451000.2014.958088.
10. Uiblein, F. & Causse, R. (2013): A new deep-water goatfish of the genus Upeneus (Mullidae) from Vanuatu, South Pacific. Zootaxa, 3666 (3): 337–344.