Fir

Firs Temporal range: 49–0 Ma PreЄ Є O S D C P T J K Pg N [1]
Korean fir (Abies koreana) cone and foliage
Scientific classification
Kingdom:	Plantae
Division:	Pinophyta
Class:	Pinopsida
Order:	Pinales
Family:	Pinaceae
Genus:	Abies Mill.
Species
See text

Firs (Abies) are a genus of 48–56 species of evergreen coniferous trees in the family Pinaceae. They are found through much of North and Central America, Europe, Asia, and North Africa, occurring in mountains over most of the range. Firs are most closely related to the genus Cedrus (cedar). Douglas firs are not true firs, being of the genus Pseudotsuga.

They are large trees, reaching heights of 10–80 m (33–262 ft) tall and trunk diameters of 0.5–4 m (1 ft 8 in–13 ft 1 in) when mature. Firs can be distinguished from other members of the pine family by the unique attachment of their needle-like leaves and by their different cones.

Identification of the different species is based on the size and arrangement of the leaves, the size and shape of the cones, and whether the bract scales of the cones are long and exserted, or short and hidden inside the cone.

Leaves

A. alba - the underside of leaves with 2 whitish strips formed by wax-covered stomatal bands

A. grandis foliage - upper side of the leaves

Atypical A. alba foliage from Dinaric calcareous fir forests on Mt. Orjen

Firs can be distinguished from other members of the pine family by the unique attachment of their needle-like leaves to the twig by a base that resembles a small suction cup.

The leaves are significantly flattened, sometimes even looking like they are pressed, as in A. sibirica.

The leaves have two whitish strips on the bottom, each of which is formed by wax-covered stomatal bands. The upper surface of the leaves usually is uniformly green and shiny, without stomata or only with few at their tips, visible as whitish spots. Some of the species however have the upper surface of leaves dull, gray-green, bluish-gray to silvery, coated by wax with variable number of stomatal bands, and not always continuous. An example species with shiny green leaves is A. alba, and an example species with dull waxy leaves is A. concolor.

The tips of leaves are usually more or less notched (as in A. firma), but sometimes rounded or dull (as in A. concolor, A. magnifica) or sharp and prickly (as in A. bracteata, A. cephalonica, A. holophylla). The leaves of young plants are usually sharper.

The way they spread from the shoot is very diverse, only in some species comb-shaped, with the leaves arranged on two sides, flat (A. alba) ^[2]

Cones

Intact and disintegrated Bulgarian fir cones

Immature cones of some of species or races are green, not purple-bluish. Manchurian fir.

Disintegrating cones of Manchurian fir

Firs differ from other conifers in having erect, cylindrical cones 5–25 cm (2.0–9.8 in) long that disintegrate at maturity to release the winged seeds.

In contrast to spruces, even large fir cones do not hang, but are raised like candles.

Mature cones are usually brown, young in summer can be green, for example:

A. grandis, A. holophylla, A. nordmanniana

or purple and blue, sometimes very dark:

A. fraseri, A. homolepis (var. umbellata green), A. koreana ('Flava' green), A. lasiocarpa, A. nephrolepis (f. chlorocarpa green), A. sibirica, A. veitchii (var. olivacea green).^[2]

Classification

• Section Abies (central, south & east Europe, Asia Minor)
  • Abies alba—silver fir
  • Abies nebrodensis—Sicilian fir
  • Abies borisii-regis—Bulgarian fir
  • Abies cephalonica—Greek fir
  • Abies nordmanniana—Nordmann fir or Caucasian fir
    • Abies nordmanniana subsp. equi-trojani—Kazdağı fir, Turkish fir
    • Abies nordmanniana subsp. bornmülleriana—Uludağ fir
  • Abies pinsapo—Spanish fir
    • Abies pinsapo var. marocana—Moroccan fir
  • Abies numidica—Algerian fir
  • Abies cilicica—Syrian fir
• Section Balsamea (Taiga|boreal Asia and North America, and high mountains further south)
  • Abies fraseri—Fraser fir
  • Abies balsamea—balsam fir
    • Abies balsamea var. phanerolepis—bracted balsam fir
  • Abies lasiocarpa—subalpine fir
    • Abies lasiocarpa var. arizonica—corkbark fir
    • Abies lasiocarpa var. bifolia—Rocky Mountains subalpine fir
  • Abies sibirica—Siberian fir
    • Abies sibirica var. semenovii—
  • Abies sachalinensis—Sakhalin fir
  • Abies koreana—Korean fir
  • Abies nephrolepis—Khinghan fir
  • Abies veitchii—Veitch's fir
    • Abies veitchii var. sikokiana—Shikoku fir
• Section Grandis (western North America to Mexico, Guatemala, Honduras and El Salvador, lowlands in north, moderate altitudes in south)
  • Abies grandis—grand fir or giant fir
    • Abies grandis var. idahoensis—interior grand fir or giant fir
  • Abies concolor—white fir
    • Abies concolor subsp. lowiana—Low's white fir
  • Abies durangensis—Durango fir
    • Abies durangensis var. coahuilensis—Coahuila fir
  • Abies flinckii—Jalisco fir
  • Abies guatemalensis—Guatemalan fir
• Section Momi (east & central Asia, Himalaya, generally at low to moderate altitudes)
  • Abies kawakamii—Taiwan fir
  • Abies homolepis—Nikko fir
  • Abies recurvata—Min fir
    • Abies recurvata var. ernestii—Min fir
  • Abies firma—Momi fir
  • Abies beshanzuensis—Baishanzu fir
  • Abies holophylla—Manchurian fir
  • Abies chensiensis—Shensi fir
    • Abies chensiensis subsp. salouenensis—Salween fir
  • Abies pindrow—Pindrow fir
  • Abies ziyuanensis—Ziyuan fir
• Section Amabilis (Pacific coast mountains, North America and Japan, in high rainfall mountains)
  • Abies amabilis—Pacific silver fir
  • Abies mariesii—Maries' fir

A. fabri, Sichuan, China

• Section Pseudopicea (Sino-Himalayan mountains, at high altitude)
  • Abies delavayi—Delavay's fir
    • Abies delavayi var. nukiangensis—
    • Abies delavayi var. motuoensis—
    • Abies delavayi subsp. fansipanensis—
  • Abies fabri—Faber's fir
    • Abies fabri subsp. minensis—
  • Abies forrestii—Forrest's fir
  • Abies densa—Bhutan fir
  • Abies spectabilis—East Himalayan fir
  • Abies fargesii— Farges' fir
  • Abies fanjingshanensis—Fanjingshan fir
  • Abies yuanbaoshanensis—Yuanbaoshan fir
  • Abies squamata—flaky fir
• Section Oiamel (Central Mexico, at high altitude)
  • Abies religiosa—sacred fir
  • Abies hickelii—Hickel's fir
    • Abies hickelii var. oaxacana—Oaxaca fir

A. magnifica, California, USA

• Section Nobilis (western U.S., high altitudes)
  • Abies procera—noble fir
  • Abies magnifica—red fir
    • Abies magnifica var. shastensis—Shasta red fir
• Section Bracteata (California coast)
  • Abies bracteata—bristlecone fir
• Section Incertae sedis
  • Abies milleri—(Extinct) Early Eocene^[1]

Uses and ecology

Wood of most firs is considered unsuitable for general timber use, and is often used as pulp or for the manufacture of plywood and rough timber. Because this genus has no insect or decay resistance qualities after logging, it is generally recommended for construction purposes as indoor use only (e.g. indoor drywall on framing). This wood left outside cannot be expected to last more than 12 to 18 months, depending on the type of climate it is exposed to. It is commonly referred to by several different names, including North American timber, SPF (spruce, pine, fir) and whitewood.

Nordmann fir, noble fir, Fraser fir and balsam fir are popular Christmas trees, generally considered to be the best for this purpose, with aromatic foliage that does not shed many needles on drying out. Many are also decorative garden trees, notably Korean fir and Fraser fir, which produce brightly coloured cones even when very young, still only 1–2 m (3.3–6.6 ft) tall. Other firs can grow anywhere between 30 and 236 feet (9.1 and 71.9 m) tall. Fir Tree Appreciation Day is June 18.

Firs are used as food plants by the caterpillars of some Lepidoptera species, including Chionodes abella (recorded on white fir), autumnal moth, conifer swift (a pest of balsam fir), the engrailed, grey pug, mottled umber, pine beauty and the tortrix moths Cydia illutana (whose caterpillars are recorded to feed on European silver fir cone scales) and C. duplicana (on European silver fir bark around injuries or canker).

Abies spectabilis or Talispatra is used in Ayurveda as an antitussive drug.^[3][4]

References

1. Schorn, Howard; Wehr, Wesley (1986). "Abies milleri, sp. nov., from the Middle Eocene Klondike Mountain Formation, Republic, Ferry County, Washington". Burke Museum Contributions in Anthropology and Natural History 1: 1–7.
2. Seneta, Włodzimierz (1981). Drzewa i krzewy iglaste (Coniferous trees and shrubs) (in Polish) (1st ed.). Warsaw: Państwowe Wydawnictwo Naukowe (PWN). ISBN 83-01-01663-9.
3. Schar (2015). "Douglas Fir, Pseudotsuga menziesii". Archives. Doctor Schar. Retrieved 2015-10-04.
4. Kershaw, Linda (2000). Edible and Medicinal Plants of the Rockies. Edmonton, AB: Lone Pine Publishing. p. 26. ISBN 978-155105-229-8.

Bibliography

Philips, Roger. Trees of North America and Europe, Random House, Inc., New York ISBN 0-394-50259-0, 1979.

External links

• Abies at The Gymnosperm Database
• Michael P. FRANKIS CONE COLLECTION: Abies at the Arboretum de Villardebelle—images of cones of selected species

^[1]

1. Platt, Karen "Gold Fever" provides descriptions of golden or yellow-leaved Abies cultivars http://www.karenplatt.co.uk